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Re: Tyrannosaurs



Here's a splitter's cladogram of Tyrannosauria I published in
_Dino-Frontline_ #9 and 10, with a mistake or two corrected. Tyrannosauria is
defined as a monophyletic suborder that includes all theropods with a
didactyl manus. I expect disagreement with this, since the differences among
the tyrannosaurids are subtle and in some workers' opinions insufficient for
generic separation--not to mention the not-as-blue-sky-as-it-at-first-seems
inclusion of _Compsognathus_ and didactyl-manus dino-birds. Numbered nodes
are "branching" nodes--cladogenetic events, if you like; unnumbered nodes are
monophyletic taxa not further split in the analysis.

1: Tyrannosauria
  Dino-birds with didactyl wings
  2: Unnamed node (ground-dwelling, cursorial forms)
    Compsognathidae
      3: Compsognathus
        Compsognathus longipes
        Compsognathus corallestris
    4: Unnamed node (forms with "tyrannosaur"-like braincase)
      Itemiridae
        Itemirus
          Itemirus medullaris
      5: Tyrannosauridae
        6: Shanshanosaurinae
          Stygivenator
            Stygivenator molnari
          7: Unnamed node
            Shanshanosaurus
              Shanshanosaurus huoyanshanensis
            8: Unnamed node
              Alectrosaurus
                Alectrosaurus olseni
              OMNH 10131 "Aublysodon" (unnamed genus/species)
        9: Tyrannosaurinae
          Alioramini
            Alioramus
              Alioramus remotus
          10: Tarbosaurini
            11: Unnamed node ("monophyletic Tarbosaurini")
              Maleevosaurus
                Maleevosaurus novojilovi
              12: Unnamed node ("big tarbosaurinids")
                Tarbosaurus
                  Tarbosaurus efremovi
                Jenghizkhan
                  Jenghizkhan bataar
            13: Albertosaurini (tyrannosaurines w/lacrimal horns)
              FMNH PR308 albertosaurinid (unnamed genus/species)
              14: Unnamed node
                15: Albertosaurus
                  UA 10 (unnamed species)
                  Albertosaurus sarcophagus
                16: Unnamed node
                  Gorgosaurus
                    Gorgosaurus libratus
                  17: Unnamed node
                    Daspletosaurus
                      Daspletosaurus torosus
                    18: Unnamed node
                      Two Medicine albertosaurinid
                      19: Tyrannosaurini
                        20: Unnamed node
                          Dinotyrannus
                            Dinotyrannus megagracilis
                          Nanotyrannus
                            Nanotyrannus lancensis
                        21: Tyrannosaurus
                          AMNH 5027 (unnamed species)
                          Tyrannosaurus rex

This looks more complicated than it is; but I can't transmit the actual
graphic via e-mail. So typographical indenting conventions will have to do.
OMNH = Oklahoma Museum of Natural History; FMNH = Field Museum of Natural
History; UA = University of Alberta; AMNH = American Museum of Natural
History

G.O.