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THIS IS A LONG ONE!!!!
Here it is at last. The "ecology" and taxonomy will probably get
altered, but the first block of info and the publications is about as
complete as I could get. I pity you having to type it in.
This is the first entry for the Dino Pages, the web-site that I am
assisting Doug McLemore in putting together. It am putting it here for
editorial reveiw. If there is ANYTHING you consider innacurrate or could
be stated better or more objectively, please say so. These entries will
provide people with a professional rather than popular interest in dinosaurs
with technical info, especially a comprehensive list of publications relating
to dinosaur genera (saving considerable trouble in discussing the genra in
greater detail). Most of the refernces come from DINOSAURIA, with the
post-1990 ones having been sent to me (thanks again, everybody) and
gleaned from Sereno and Nova's papers. These are mostly technical
publications, and there is a Discover article in there I'm not really sure
belongs. There is also and article by Olsen and Sues I don't know the
Doug has set up a taxonomic guide for the Dino Pages based on the
the one Lessom and Glut used in Dinosaur Encyclopedia. Recognizing the
unsettled nature of dinosaur taxonomy, we are toying around with the idea of
having three or four popular alternate phylogenies to look at and use to
find your way around genera. If you want to be extremely accomidating, post
complete dinosaur family trees (only down to the family level, I'm a
reasonably man) that are popular and commonly accepted. I'd really like
to have G.Os too, if he doesn't mind (that wasn't meant like it
sounded). Its a pity no one ever set up a web site devoted entirely to
AND FINALLY: anyone care to explain internet policy on scanning
other peoples artwork onto web sites? I'd like to be polite and ask
permission before submitting Doug a restoration or skeletal recontruction,
particularly if the artist is likely to visit the web-site.
GENUS _Herrerasaurus_ (Reig, 1963)
_Herrerasaurus_ _ischigualastensis_ (Reig, 1963)
Etymology: "Herrera's lizard" for Victorino Herrera, finder of the holotype
"from ischigualasto" after the Ischigualasto formation
Synonyms: ?_Ischisaurus_ _cattoi_ (Reig, 1963)
?_Frenguellisaurus_ _ischigualastensis_ (Novas, 1986)
?cf. _Staurikosaurus_ (Brinkman and Sues, 1987)
Holotype: PVL 2566
Referred material: MACN 18.060 (holotype: _I.cattoi_), MACN 18.688, MCZ
7064, MLP 61-VIII-2-2 (I.cattoi), PVL 2054, PVL 2558a, PVSJ 53 (holotype:
_F.ischigualastensis_), PVSJ 104, PVSJ 373, PVSJ 407, PVSJ 461
Formation & location: Lower Ischigualasto Formation
San Juan, Argentina
Age: Carnian (225 MYBP)
(Doug, this is from the Sereno & Novas papers and is disputed by some.
Incorporate it or ditch it or set it aside for the time
being as you see fit, depending on how you decide to handle taxonomy.)
THEROPODA (Marsh, 1881)
HERRERASAURIDAE (Bendetto, 1973)
Sereno and Novas (1992) consider _Herrerasaurus_ to be not only a
true saurischian, but a basal theropod as well. Novas (1993) places all
carniverous dinosaur material from the Ischigualasto formation into this
_Herrerasaurus_ _ischigualastensis_ was one of the earliest
dinosaurs, and the largest bipedal carnivore that had yet to appear. Its
lower jaw had a hinge joint which may have facilitated a more gripping
bite. The hands seem well adapted for grasping prey.
The early ornithischian _Pisanosaurus_ _mertii_, as well as
rynchosaurs and other synapsids are also known from the Ischigualasto
Formation and may have been primary prey items. _H.ischigualastensis_
may have faced competition and possibly predation from the larger and
more abundant rauisuchian thecodonts of the time.
Facies and plant fossils from the Ischigualasto Formation seem to
indicate a seasonal climate upland forest of sphenopsids, ferns, and seed
Bendetto, J.L. 1973. Herrerasauridae, neuva familia de saurisquios
Triasicos. Ameghiniana 10: 89-102
Bonaparte, J.F. 1982. Faunal replacement in the Triassic of South
America. J. Vertebr. Paleontol. 2: 362-371
Brinkman, D.B. and Sues, H.-D. 1987. A staurikosaurid dinosaur from the
Upper Triassic Ischigualasto Formation of Argentina and the relationships
of the Staurikosauridae. Paleontology 30: 493-503
Galton, P.M. 1985. The poposaurid thecodontian _Teratosaurus_ _suevicus_
v. Meyer, plus referred specimens mostly based on prosauropod dinosaurs,
from the Middle Stubensandstein (Upper Triassic) of Nordwurttemberg.
Stuttgarter Beitr. Naturk. B 118: 1-29
Kluger, J. 1990. Tripping over history. Discover 11(12): 40-41
Novas, F.E. 1989a. The tibia and tarsus in Herrerasauridae (Dinosauria,
insertae sedis) and the origin and evolution of the dinosaur tarsus. J.
Paleontol. 63: 677-690
---- 1989b. Los dinosaurios carnivoros de la Argentina. Ph.D.
dissertation (unpublished), Universidad Nacional de La Plata, La Plata,
---- 1992. Phylogenetic relationships of the basal dinosaurs, the
Herrerasauridae. Paleontology 35: 51-62
---- 1993. New information on the systematics and postcranial skeleton of
_Herrerasaurus_ _ischigualastensis_ (Theropoda: Herrerasauridae) from
the Ischigualasto Formation (Upper Triassic) of Argentina. J.
Vertebr. Paleontol. 13(4): 400-423
Olsen, P.E. and Sues, H.-D. 1986. pp. 321-351 in K. Padian (ed.), The
Beginning of the Age of Dinosaurs. Cambridge University Press, Cambridge
Paul, G. 1988. Predatory Dinosaurs of the World. A Complete Illustrated
Guide. Simon and Schuster, New York, 464 pp.
Reig, O.A. 1963. La presencia de dinosaurios saurisquios en los "Estratos
de Ischigualasto" (Mesotriassico superior) de las Provincias de San
Juan y La Rioja (Republica Argentina). Ameghiniana 3: 3-20
Sereno, P.C. 1993. The pectoral girdle and forelimb of the basal theropod
_Herrerasaurus_ _ischigualastensis_. J. Vertebr. Paleontol. 13(4):
Sereno, P.C. and Novas, F.E. 1990. Dinosaur origins and the phylogenetic
position of pterosaurs. J. Vertebr. Paleontol. 10: 42A
---- 1992. The complete skull and skeleton of an early dinosaur. Science
---- 1993. The skull and neck of the basal theropod _Herrerasaurus_
_ischigualastensis_. J. Vertebr. Paleontol. 13(4): 451-476
Sues, H.-D. 1990. Staurikosauridae and Herrerasauridae; pp. 143-147 in
D.B. Weishampel, P. Dodson, and H. Osmolska (eds.), The Dinosauria.
University of California Press, Berkely and Los Angeles
Wellnhofer, P. 1993. News on the evolution of dinosaurs.
Naturwissenschaftliche Rundschau 46(12): 489 (in German)