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Question about megatracksites

Hello all,

I am reading Lockley's "Tracking Dinosaurs" and in chapter 4, page 44 it says:

 "[certain large tracksites] are often associated with the boundary layers 
between geological formations"

Can someone explain to me why this is so? I suppose it has to do with... 
well... I don't know, that's why I ask.

Cheers, happy new year and all.