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Dinosaur Genera List update #198



January, 2003 has seen the addition of no less than five names to the 
Dinosaur Genera List. Two appear to be the first two new genera for 2003, one 
appears to be the oldest generic name applied to a dinosaur fossil, and the 
last two are two more previously unpublished synonyms for an already named 
British dinosaur, Eotyrannus. I have duly recorded these in The Dinosaur 
Catalogue and I'll run their Catalogue entries here as they currently stand.

The latest issue of the Journal of Vertebrate Paleontology introduces two new 
dinosaur genera and three new species, so we add these two names as #966 and 
967 respectively:

Heyuannia Lü, 2003
Sphaerotholus Williamson & Carr, 2003

The citations are, respectively:

Lü Junchang, 2003. "A new oviraptorosaurid [sic](Theropoda: Oviraptorosauria) 
from the Late Cretaceous of southern China," Journal of Vertebrate 
Paleontology 22(4): 871?875 [December 2002 issue, published January 14, 2003].

Williamson, Thomas E. & Carr, Thomas D., 2003. "A new genus of derived 
pachycephalosaurian from western North America," Journal of Vertebrate 
Paleontology 22(4): 779?801 [December 2002 issue, published January 14, 2003].

And these are The Dinosaur Catalogue entries for these genera as they 
currently stand (apologies for the inevitable loss of typographical 
information for the entries due to email and listservs; additions and 
corrections welcomed). Note that two species were described for the genus 
Sphaerotholus, and that the Heyuannia paper carried a misspelling of the 
generic name that also gets its own minor Catalogue entry:

    Heyuania Lü, 2003: Misspelling of Heyuannia
Heyuania huangi Lü, 2003
    Misspelling of Heyuannia huangi

    Heyuannia Lü, 2003
    Heyuania Lü, 2003: Misspelling
    Theropoda > Tetanurae > Coeluria > Maniraptora > Oviraptorosauria > 
Oviraptoridae
Heyuannia huangi Lü, 2003?
    Heyuania huangi Lü, 2003: Misspelling in original description
    Late Cretaceous > Maastrichtian(?) > Dalangshan Fm.
    Asia > China >Guangdong Prov. > Heyuan City > quarry near Huangsha village
    HYMV1-1: Partial articulated skeleton w/ skull material, missing only 
forelimbs and distal caudal vertebrae
    Hypodigm APK:25/75: HYMV1-2 furcula, partial right shoulder girdle and 
forelimb; HYMV1-3 partial right manus, and HYMV1-4, both associated with 
holotype skeleton; and MYMV1-5, left manus; all in addition to holotype 
skeleton and likely belonging to the same individual

        Sphaerotholus Williamson & Carr, 2003
    Ornithischia > Marginocephalia > Pachycephalosauria >
    Pachycephalosauridae > Pachycephalosaurinae
Sphaerotholus buchholtzae Williamson & Carr, 2003
    Upper Cretaceous > Upper Maastrichtian > Hell Creek Fm.
    North America > United States > Montana > Carter Co. > locality SE¼, Sec. 
5, T1S R55E
    TMP 87.113.3: Skull dome
    Hypodigm APK:40/0: Holotype partial skull only, originally referred to 
the species Stegoceras  edmontonense (Giffin, 1989) and thence to 
Prenocephale edmontonensis (Sullivan, 2000)

Sphaerotholus goodwini Williamson & Carr, 2003?
    Upper Cretaceous > Upper Campanian > Kirtland Shale > De-na-zin Mbr. (= 
"upper shale member")
    North America > United States > New Mexico > San Juan Basin > San Juan 
Co. > Bisti/De-na-zin Wilderness Area > Willow Wash > NMMNH locality L-3921
    NMMNH P-27403: Skull dome lacking facial and palatal bones
    Hypodigm APK:50/0: NMMNH P-30068 nearly complete left dentary w/ 
squamosal and cranial fragments, in addition to holotype skull dome

The Sphaerotholus paper carries additional taxonomic information about 
pachycephalosaurians that will be included in other entries in The Dinosaur 
Catalogue. I originally entered these names into the Dinosaur Genera List 
with the dates 2002, because December 2002 is the cover date of the JVP, but 
once I received the issue, I saw that its publication date was January 14, 
2003, so I changed the year of publication to 2003.

------------------------------------------------------------

The fourth new dinosaur species for 2003 appeared in the January 23, 2003 
issue of Nature. It doesn't impact the Dinosaur Genera List, since it is a 
second species of the genus Microraptor, already in the List, but it inspired 
me to complete the Microraptor entry for The Dinosaur Catalogue:

        Microraptor Xu, Zhou & Wang, 2000
    Junior objective synonym of rejected (pending) generic name Archaeoraptor
    Possible senior subjective synonym of Cryptovolans
    Theropoda > Tetanurae > Coeluria > Maniraptora > Deinonychosauria > 
Dromaeosauridae
Microraptor gui Xu, Zhou, Wang, Kuang, Zhang & Du, 2003
    Possible junior subjective synonym of Cryptovolans pauli
    Lower Cretaceous > Jehol Gr. > Upper Jiufotang Fm.
    Asia > China > Chaoyang Basin > Liaoning Prov. > Chaoyang Co. > 30 km SW 
of Chaoyang City > Dapingfang
    IVPP V13352: Nearly complete skeleton in slab and counterslab, missing 
anterior portion of skull, showing detailed feather impressions 
    Hypodigm APK:50/100: IVPP V13320 nearly complete skeleton w/ feather 
impressions, referred to this species in original description; also used in 
the original description were partial skeletons with feather impressions 
TNP00996,  IVPP V13351, and IVPP V13476 (as Microraptor sp.), and IVPP V13477 
(as Dromaeosauridae sp. indet.), the latter two from Shangheshou (3 km NW of 
Chaoyang City)

Microraptor zhaoianus Xu, Zhou & Wang, 2000?
    Junior objective synonym of rejected (pending) name Archaeoraptor 
liaoningensis
    Lower Cretaceous > Jehol Gr. > Upper Jiufotang Fm.
    Asia > China > Chaoyang Basin > Liaoning Prov. > Chaoyang Co. > 
Xiasanjiazi
    IVPP V12330: Complete skeleton in slab and counterslab, showing feather 
impressions
    Hypodigm APK:100/100: Holotype skeleton only; tail vertebrae from the 
IVPP V12330 counterslab were made the lectotype of Archaeoraptor liaoningensis
 by Olson, 2000 (cf.), after it was discovered that they had been improperly 
integrated into a slab with the holotype skeleton of Archaeovolans repatriatus
 (cf.) but before IVPP V12330 had been made the holotype of Microraptor 
zhaoianus; a petition to the ICZN to suppress Archaeoraptor and Archaeoraptor 
liaoningensis in favor of Microraptor and Microraptor zhaoianus is in 
preparation (Olshevsky, 2001)

I haven't yet had time to prepare that ICZN petition referred to above, but I 
hope to have it finished later this year. Microraptor gui is, of course, the 
four-winged dino-bird that has received so much publicity lately.

Xu Xing, Zhou Zhonghe, Wang Xiaolin, Kuang Xuewen, Zhang Fucheng & Du 
Xiangke, 2003. "Four-winged dinosaurs from China," Nature 421(6921): 335?340 
[January 23, 2003].

Prum, Richard O., 2003. "Dinosaurs take to the air," Nature 421(6921): 
323?324 [January 23, 2003].

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In a most interesting paper on the history of dinosaur discoveries (thanks to 
Tracy Ford for finding this one), Delair & Sarjeant report a dinosaur fossil 
originally described by Edward Lhuyd (or Edvardus Lhuyd) in 1699 and actually 
given a binominal nearly 60 years before Linnaeus introduced the current 
system of biological nomenclature. While the date precludes acceptance of the 
name as scientifically valid, it nevertheless qualifies for inclusion in the 
Dinosaur Genera List, and supersedes Scrotum humanum Brookes, 1763 as the 
oldest name applied to a dinosaur fossil in paleontological literature. 
Accordingly, I add as name #968

Rutellum Lhuyd, 1699 [nomen oblitum; pre-Linnaean name: cf. Delair & 
Sarjeant, 2002]

Here is The Dinosaur Catalogue entry:

    Rutellum Lhuyd, 1699 [nomen oblitum]
    Brontosauria > Sauropoda > Cetiosauridae
Rutellum implicatum Lhuyd, 1699?
    Jurassic > Thames gravels or Coral Rag
    Europe > Great Britain > England > Oxfordshire > 8 km SW of Whitney > 
Carswell
    Lhuyd, 1699 specimen #1352 (University of Oxford collection, lost)
    Hypodigm APK: "Type" tooth only, as reported and illustrated by Delair & 
Sarjeant, 2002, identified as a cetiosaurid tooth
    This is the oldest species binominal of a dinosaur thus far discovered in 
the literature, preceding by nearly 60 years the origin of the Linnaean 
system and therefore not a scientifically valid name; catalogued here as a 
nomen oblitum (forgotten name)

Delair, Justin B. & Sarjeant, William A. S., 2002. "The earliest discoveries 
of dinosaurs; the records re-examined," Proceedings of the Geologists' 
Association 113: 185?197.

Lhuyd, E., 1699. Lithophylacii Britannici Ichnographia, sive, lapidium 
aliorumque fossilium Britannicorum singulari figura insignium, Gleditsch & 
Weidmann, London [pagination not available].

The paleontological community laments the recent passing of Bill Sarjeant, a 
nonpareil historian of dinosaurology and dinosaur ichnologist. The paper with 
Justin Delair was published posthumously.

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Finally, Darren Naish informs me of two more published junior synonyms of 
Eotyrannus, which become names #969 and 970 in the Dinosaur Genera List:

Kittysaurus Hargreaves, 2001 [nomen nudum -> Eotyrannus]
Fusinasus Hutt, 2002 [nomen nudum -> Eotyrannus]

The citations are:

Hargreaves, Richard, 2001. "He's daddy of the dinosaurs," The News, May 10, 
2001: 13.

Hutt, S. 2002. "Mr Leng's dinosaur," The Geological Society 
of the Isle of Wight Newsletter 2(6), 12?14 [December 2002].

And The Dinosaur Catalogue entries are:

    Kittysaurus Hargreaves, 2001 [nomen nudum]
    Junior objective synonym of Eotyrannus
    Theropoda > Tetanurae > Coeluria > Tyrannosauria
Generic name revealed in a popular-science news article as an unused name 
considered for the genus Eotyrannus

    Fusinasus Hutt, 2002 [nomen nudum]
    Junior objective synonym of Eotyrannus
    Theropoda > Tetanurae > Coeluria > Tyrannosauria
Fusinasus lengi Hutt, 2002?
    Junior objective synonym of Eotyrannus lengi
    Generic and specific names revealed in a popular-science news article as 
having once been considered for the genus and species Eotyrannus lengi

The Dinosaur Genera List is at website

http://members.aol.com/Dinogeorge/dinolist.html