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Kellner, Alexander W.A., Aguirre-Urreta, María B. and Ramos, Victor A. On the pterosaur remains from the Río Belgrano formation (Barremian), Patagonian Andes of Argentina. An. Acad. Bras. Ciênc., Dec 2003, vol.75, no.4, p.487-495.

Pterosaur remains from the Río Belgrano Formation, Santa Cruz Province, Argentina, were found close to the Estancia Río Roble, along with several ammonoids that indicate a Barremian age for those strata. The specimens (MACN-SC 3617) consist of one ulna and one element tentatively identified as a portion of a wing metacarpal. The ulna shows morphological affinities with the Pteranodontoidea (sensu Kellner 1996), particularly with the members of the Anhangueridae by having a well developed ventral crest close to the proximal articulation, and is tentatively referred to this pterosaur clade. The oldest record of the Anhangueridae, previously limited to the Aptian/Albian, is therefore extended to the Barremian. The Argentinean material is preserved in three dimensions, an unusual condition for pterosaur fossils from that country, indicating that the site situated near the Estancia Río Roble has a great potential for new and well preserved specimens.

Available on: http://www.scielo.br/pdf/aabc/v75n4/a07v75n4.pdf