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re: Anhangueridae/Ornithocheiridae

Three questions:

1. Was wondering where Haopterus, Arthurdactylus and Istiodactylus fit
in the Ornithocheiridae (sensu Unwin), if anywhere?

2. Was wondering if there are any subclades within the Ornithocheiridae?

3. Was wondering which taxon is the closest known sister taxon to the

I have still not seen the paper, hence the topical questions.

David Peters
St. Louis