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Re: South American Pterosaurs



Ivan Kwan wrote-

My data is following the taxonomy of Unwin 2000....

> 1) How many species of _Anhanguera_ are recognised? _Anhanguera
> blittersdorffi_ & _Anhanguera santanae_? (The paper does mention that _A.
> santanae_ may be a juvenile _Coloborhynchus_)

Anhanguera Campos and Kellner 1985
A. blittersdorffi Campos and Kellner 1985
Aptian, Early Cretaceous
Santana Formation, Brazil
A. santanae Wellnhofer 1985
Aptian, Early Cretaceous
Santana Formation, Brazil
A. cuvieri (Bowerbank vide Seeley 1865) Unwin et al. 2000
= Pterodactylus cuvieri Bowerbank vide Seeley 1865
= Ptenodactylus "brachyrhynchus" Seeley 1869
= Ptenodactylus "colorhinus" Seeley 1869
= Ptenodactylus cuvieri Seeley 1869
= Ptenodactylus "dentatus" Seeley 1869
= Ptenodactylus "denticulatus" Seeley 1869
= Ptenodactylus "enchorhynchus" Seeley 1869
= Ptenodactylus "scaphorhynchus" Seeley 1869
= Ornithocheirus brachyrhynchus Seeley 1870
= Ornithocheirus colorhinus Seeley 1870
= Ornithocheirus cuvieri Seeley 1870
= Ornithocheirus dentatus Seeley 1870
= Ornithocheirus denticulatus Seeley 1870
= Ornithocheirus enchorhynchus Seeley 1870
= Ornithocheirus scaphorhynchus Seeley 1870
= Ornithocheirus xiphorhynchus Seeley 1870
= Coloborhynchus cuvieri Owen 1874
= Pterodactylus brachyrhynchus Sauvage 1882
= Pterodactylus colorhinus Sauvage 1882
= Pterodactylus dentatus Sauvage 1882
= Pterodactylus enchorhynchus Sauvage 1882
= Pterodactylus scaphorhynchus Sauvage 1882
= Lonchodectes scaphorhynchus Hooley 1914
Albian, Early Cretaceous
Toolebuc Formation, Queensland, Australia; Cambridge Greensand, England
A. fittoni (Owen 1859) Unwin et al. 2000
= Pterodactylus fittoni Owen 1959
= Ptenodactylus fittoni Seeley 1869
= Ptenodactylus "nasutus" Seeley 1869
= Ptenodactylus "polyodon" Seeley 1869
= Ornithocheirus fittoni Seeley 1870
= Ornithocheirus nasutus Seeley 1870
= Ornithocheirus polyodon Seeley 1870
= Pterodactylus nasutus Sauvage 1882
= Pterodactylus polydo Sauvage 1882
Albian, Early Cretaceous
Cambridge Greensand, England
A. sp.
Cenomanian, Late Cretaceous
Lysaya Gora, Russia
Material- (ZIN PNT-S50-1) dentary fragment

> 2) _Anhanguera piscator_ is now shifted into _Coloborhynchus_? Is this
> generally accepted?

That's how I have it catalogued-
Coloborhynchus robustus (Wellnhofer 1987) Fastnacht 2001
= Tropeognathus robustus Wellnhofer 1987
= Anhanguera robustus Kellner and Campos 1989
= Anhanguera piscator Kellner and Tomida 2000

> 3) Has _Tropeognathus_ been sunk into _Criorhynchus_? I noticed that
> _Tropeognathus mesembrinus_ was called _Criorhynchus mesembrinus_, and
> _Tropeognathus ribustus_ was now _Coloborhynchus robustsus_.

Ornithocheirus Seeley 1869
= Criorhynchus Owen 1874
= Tropeognathus Wellnhofer 1987
O. simus (Owen, 1861) Seeley 1869
= Pterodactylus simus Owen 1861
= Pterodactylus "carteri" Seeley 1865
= Ornithocheirus "carteri" Seeley 1869
= Ornithocheirus "platyrhinus" Seeley 1869
= Ornithocheirus carteri Seeley 1870
= Ornithocheirus platyrhinus Seeley 1870
= Criorhynchus simus Owen 1874
= Criorhynchus carteri Hooley 1914
= Criorhynchus platyrhinus Hooley 1914
Albian, Early Cretaceous
Cambridge Greensand, England
O. mesembrinus (Wellnhofer 1987)
= Tropeognathus mesembrinus Wellnhofer 1987
O. sp. (Unwin, 2000)
Albian Early Cretaceous
Upper Greensand, England

> 4) Which species of _Ornithocheirus_ are recognised?

Besides the valid ones above-

Pterosauria indet.

Ornithocheirus? clifti (Mantell 1835) Newton 1888
= Palaeornis Mantell 1835 (preoccupied Vigors 1825)
= "Osteornis" Gervais 1844
= "Lithosteornis" Gervais 1844
= Palaeornis clifti Mantell 1835
= "Osteornis ardeaceus" Gervais 1844
= "Lithosteornis ardeaceus" Gervais 1844
= Pterodactylus silvestris Owen 1845
= Pterodactylus clifti Bronn 1848
Early Cretaceous, England
Holotype- (BMNH 02353) partial humerus

Ornithocheirus? curtus (Owen 1870) Newton 1888
= Pterodactylus curtus Owen 1870
Early Cretaceous
Wealden, England
Holotype- (BMNH R1440) distal metacarpal IV

Ornithocheirus? diomedeus (Owen 1846) Newton 1888
= "Osteornis diomedeus" Owen 1844
= Cimoliornis Owen 1846
= Cimoliornis diomedeus Owen 1846
= Pterodactylus diomedeus Owen 1851
= Pterodactylus giganteus Morris 1854 (non Oken 1819, non Bowerbank 1846)
Late Cretaceous
England
Holotype- (BMNH 39418) distal metacarpal

Ornithocheirus? "macrorhinus" Jukes-Browne 1875
= Ptenodactylus "macrorhinus" Seeley 1869
= Pterodactylus "macrorhinus" Sauvage 1882
Albian, Early Cretaceous
Cambridge Greensand, England

Ornithocheirus? "oxyrhinus" Owen 1870
= Ptenodactylus "oxyrhinus" Seeley 1869
= Pterodactylus "oxyrhinus" Sauvage 1882
Albian, Early Cretaceous
Cambridge Greensand, England

Ornithocheirus? weidenrothi Wild 1990
Early Cretaceous
Germany

Azhdarchidae

Ornithocheirus? bunzeli Seeley 1881
Maastrichtian, Late Cretaceous
Gosau Formation, Austria
Comments- Possibly azhdarchid (Unwin pers. comm. to Nessov 1991)

> 7) I now suppose that _Coloborhynchus_ has 4 species: _C. piscator_, _C.
> robustus_, _C. spielbergi, C. araripensis_.

Coloborhynchus Owen 1874
= Ptenodactylus Seeley 1869 (preoccupied Gray 1845)
= Amblydectes Hooley 1914
= Siroccopteryx Mader and Kellner 1999
C. clavirostris Owen 1874
Valanginian, Early Cretaceous
Hastings Beds, England
Holotype- (BMNH R1822) anterior snout
C. sedgwickii (Owen 1859) Owen 1874
= Pterodactylus sedgwickii Owen 1859
= Pterodactylus woodwardi Owen 1861
= Ptenodactylus "crassidens" Seeley 1869
= Ptenodactylus sedgwickii Seeley 1869
= Ptenodactylus woodwardi Seeley 1869
= Ornithocheirus crassidens Seeley 1870
= Ornithocheirus sedgwickii Seeley 1870
= Ornithocheirus woodwardi Seeley 1870
= Criorhynchus woodwardi Owen 1874
= Pterodactylus crassidens Sauvage 1882
= Amblydectes crassidens Hooley 1914
= Criorhynchus crassidens Wellnhofer 1978
Albian, Early Cretaceous
Cambridge Greensand, England
C. capito (Seeley 1870) Unwin et al. 2000
= Ptenodactylus "capito" Seeley 1869
= Ptenodactylus "euryganthus" Seeley 1869
= Ornithocheirus capito Seeley 1870
= Ornithocheirus eurygnathus Seeley 1870
= Ornithocheirus reedi Seeley 1870
= Amblydectes eurygnathus Hooley 1914
= Criorhynchus capito Hooley 1914
= Criorhynchus reedi Hooley 1914
Comments- possibly male C. sedgwickii.
Albian, Early Cretaceous
Cambridge Greensand, England
C. robustus (Wellnhofer 1987) Fastnacht 2001
= Tropeognathus robustus Wellnhofer 1987
= Anhanguera robustus Kellner and Campos 1989
= Anhanguera piscator Kellner and Tomida 2000
Aptian, Early Cretaceous
Santana Formation, Brazil
References- Kellner and Tomida, 2000. Description of a new species of
Anhangueridae (Pterodactyloidea) with comments on the pterosaur fauna from
the Santana Formation (Aptian-Albian), northeastern
Brazil. National Science Museum Monographs 17, 1-135. [National Science
Museum, Tokyo, ISSN 1342-9574].
Fastnacht, 2001. First record of Coloborhynchus (Pterosauria) from the
Santana Formation (Lower Cretaceous) of the Chapada do Araripe, Brazil.
Palaontologische Zeitschrift 75, 23-36.
C. spielbergi Veldmeijer 2003
Albian, Early Cretaceous
Romualdo Member of Santana Formation, Brazil
Holotype- (RGM 401 880) skull, mandible, ceratobranchial fragments, seventh
and eighth cervical vertebrae, notarium, seven dorsal vertebrae, five rib
fragments, scapulocoracoids, humerus, ulna, radius, syncarpal, manual
phalanx, three manual unguals, pelvis, femora, tibiae
Reference- Veldmeijer, 2003. Coloborhynchus spielbergi sp. nov.
(Pterodactyloidea) from the Albian (Lower Cretaceous) of Brazil. Scripta
Geologica. 125 35-139.
C. moroccensis (Mader and Kellner 1999) Unwin 2001
= Siroccopteryx moroccensis Mader and Kellner 1999
Albian-Cenomanian, Early Cretaceous
Ksar es Souk, Morocco
Holotype- (LINHM 016) anterior snout
Reference- Mader, B. J. and Kellner, A. W. A. 1999. A new anhanguerid
pterosaur from the Cretaceous of Morocco. Boletim do Museu Nacional,
Geologia 45: 1-11.
C. wadleighi Lee 1994
Late Albian, Early Cretaceous
Paw Paw Formation, Texas, US
Holotype- (SMU 73058) anterior snout
Reference- Lee, Yuong-Nam, 1994. The Early Cretaceous Pterodactyloid
Pterosaur Coloborhynchus from North America. Paleontology, Volume 37, Part
4: 755-763.
C. sp. (Unwin and Bakhurina, 2001)
Aptian-Albian, Early Cretaceous
Zuunbayan Formation, Mongolia
Material- (MMNH 100/30) (5.5 m) anterior snout, incomplete mandible,
incomplete postcranium

> 8) Whatever happened to _Araripedactylus conandoylei_??

Two taxa, right?

Araripedactylus Wellnhofer 1977
A. dehmi Wellnhofer 1977
Aptian, Early Cretaceous
Santana Formation, Brazil

Arthurdactylus Frey and Martill 1994
A. conandoylei Frey and Martill 1994
Aptian, Early Cretaceous
Crato Formation, Brazil

Mickey Mortimer