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Tyrannosauroids and dromaeosaurs



Well, I just ran my analysis to see what it says about the recent questions
about tyrannosauroid and dromaeosaur phylogeny.

I coded Iliosuchus (9/243 characters codable) and Aviatyrannis (24/243).
Technically, I can code Aviatyrannis tentatively for a couple other
characters, but I'll need to look at Zinke (1998) at home to do that.  Of
course, I have no pubic, ischial or hindlimb characters yet.  And there are
six other ilial characters I have written down but haven't added yet.  But I
have eight more characters than last time (these damn ilial characters use
too many angles and ratios, so take foever to code), and a few added taxa
(Adasaurus, Calamosaurus, Fukuiraptor).  I also changed several cranial and
cervical characters to be MUCH less subjective.

90% majority rule consensus tree

|--+--Monolophosaurus
|  `--Sinraptor
`--+--Dryptosaurus
   `--+--Allosaurus
      `--+--"Chilantaisaurus" maortuensis
         `--+--+--Iliosuchus
            |  |--Fukuiraptor
            |  |--Stokesosaurus
            |  `--+--Bagaraatan
            |     |--Aviatyrannis
            |     `--Tyrannosauridae
            `--+--Calamosaurus
               `--+--Eotyrannus
                  `--+--Richardoestesia
                     `--+--+--Ornitholestes
                        |  `--Nqwebasaurus
                        `--+--Sinosauropteryx
                           `--+--Compsognathus
                              `--+--+--Proceratosaurus
                                 |  `--+--Coelurus
                                 |     `--+--Scipionyx
                                 |        `--YPM 1996
                                 `--+--+--+--Mononykinae
                                    |  |  `--+--Patagonykus
                                    |  |     `--Alvarezsaurus
                                    |  `--+--+--Garudimimus
                                    |     |  `--+--Archaeornithomimus
                                    |     |     `--Ornithomimidae
                                    |     `--+--Pelecanimimus
                                    |        |--"Grusimimus"
                                    |        `--+--Harpymimus
                                    |           |--Shenzhousaurus
                                    |           `--Deinocheirus
                                    `--+--+--Alxasaurus
                                       |  `--+--+--Erliansaurus
                                       |     |  `--+--Nothronychus
                                       |     |     `--+--Neimongosaurus
                                       |     |        `--Beipiaosaurus
                                       |     `--+--Erlikosaurus
                                       |        `--+--Therizinosaurus
                                       |           `--+--Nanshiungosaurus
                                       |              `--Segnosaurus
                                       `--+--Yixianosaurus
                                          `--+--+--+--Caudipteryx zoui
                                             |  |  `--Caudipteryx sp. nov.
                                             |  `--+--+--Incisivosaurus
                                             |     |  `--Avimimus
                                             |     `--+--Nomingia
                                             |        `--+--Caenagnathidae
                                             |           `--+--Microvenator
                                             |              `--Oviraptoridae
                                             `--+--Protarchaeopteryx
                                                `--+--+--Archaeornithoides
                                                   |  `--+--Byronosaurus
                                                   |     `--+--Pyroraptor
                                                   |
`--+--Sinovenator
                                                   |           `--+--IGM
100/44
                                                   |
`--+--Sinornithoides
                                                   |
`--+--Troodon
                                                   |
`--+--Saurornithoides mongoliensis
                                                   |
`--Saurornithoides junior
                                                   `--+--Scansoriopterygidae
                                                      `--+--+--Archaeopteryx
                                                         |  `--Wellnhoferia
                                                         `--Eumaniraptora

|--+--+--Microraptor gui
|  |  `--+--Microraptor zhaoianus
|  |     `--Cryptovolans
|  `--+--+--Sinornithosaurus
|     |  `--NGMC 91
|     `--+--Bambiraptor
|        `--+--Saurornitholestes
|           `--+--IGM 100/1015
|              `--+--+--Achillobator
|                 |  `--+--Utahraptor
|                 |     `--Velociraptor
|                 `--+--Deinonychus
|                    |--Dromaeosaurus
|                    `--Adasaurus
`--+--Unenlagia
   `--+--Rahonavis
      `--+--Shenzhouraptor
         `--+--Jixiangornis
            `--+--Yandangornis
               `--+--+--Sapeornis
                  |  `--Omnivoropteryx
                  `--+--+--Confuciusornis
                     |  `--Changchengornis
                     `--+--Jibeinia
                        `--+--+--Protopteryx
                           |  `--Longipteryx
                           `--+--Eocathayornis
                              |--Catalan nestling
                              |--Longirostravis
                              |--Sinornis
                              |--Concornis
                              |--Iberomesornis
                              `--+--+--Noguerornis
                                 |  |--Boluochia
                                 |  |--Neuquenornis
                                 |  |--Gobipteryx
                                 |  `--Enantiornis
                                 `--+--Eoalulavis
                                    |--+--Ambiortus
                                    |  `--Apsaravis
                                    |--+--Patagopteryx
                                    |  `--+--Liaoningornis
                                    |     `--Songlingornis
                                    `--+--Yanornis
                                       `--+--Yixianornis
                                          `--Ornithurae

Notable changes since last time
(http://www.cmnh.org/dinoarch/2003Dec/msg00549.html) -
- Yay!  Tyrannosauroids are now coelurosaurs.
- Proceratosaurus is a coelurid.
- Therizinosaurus, Nanshiungosaurus and Neimongosaurus are segnosaurs.
- Pelecanimimus is a deinocheirid.
- Sereno-esque non-avepectoran segnosaurs.
- The lone maniraptorans change position (Yixianosaurus sister to Avepectra;
Protarchaeopteryx as basalmost paravian; Pyroraptor in Troodontidae).
- Archaeornithoides is a troodontid, close to Byronosaurus.
- Troodontidae rearranged, with Byronosaurus most basal, and Saurornithoides
monophyletic.
- Paulian non-eumaniraptoran archaeopterygids, and scansoriopterygids follow
them there.
- Microraptor/Cryptovolans as basal deinonychosaurs.
- Unenlagia as a basal avialan.
- Long-tailed avialans paraphyletic.

Mickey Mortimer