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Huaxiagnathus orientalis pdf



Sorry if it has been posted. The pdf file or the Huaxiagnathus paper is 
available in the web page of the first author:

http://research.amnh.org/users/sunny/cv.html


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S. H. Hwang, M. A. Norell, Ji Q. and Gao K. 2004. A LARGE COMPSOGNATHID FROM 
THE EARLY CRETACEOUS YIXIAN FORMATION OF CHINA. Journal of Systematic 
Palaeontology, 2(1), 13-30.

Abstract

A new, large compsognathid theropod, Huaxiagnathus orientalis gen. et sp. nov., 
from the Early Cretaceous Yixian Formation deposits of Liaoning Province, 
People's Republic of China is described. The holotype specimen is nearly 
complete, lacking only the distal portion of the tail. It is the second largest 
theropod taxon discovered from Jehol Group sediments. Like all compsognathids, 
Huaxiagnathus has short forelimbs and a relatively unspecialised coelurosaur 
body plan. Previously, fairly complete skeletons existed for only two 
small-bodied taxa of compsognathids, Compsognathus longipes from the Late 
Jurassic of Western Europe and Sinosauropteryx prima, also from the Yixian. The 
phylogenetic position of Huaxiagnathus orientalis was analysed using an 
extensive matrix of theropod characters from many taxa. Huaxiagnathus 
orientalis fell out at the base of the Compsognathidae, as it lacks the 
forelimb adaptations of more derived compsognathids. The addition of 
Huaxiagnathus and t!
 he two other compsognathid species to our data matrix resulted in the 
placement of Compsognathidae near the base of Maniraptora. Furthermore, 
Alvarezsauridae, Paraves, and a monophyletic Therizinosauroidea + 
Oviraptorosauria clade fall out in an unresolved trichotomy in the strict 
consensus of our most parsimonious trees.

Key Words: Compsognathidae; Theropod; Yixian Formation; Liaoning; Jehol