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The Dinosauria 2nd Ed. - New and Excluded Theropoda



There is a LOT of information in this book the DML should know about.

First, there are three new theropod taxa, which seem to be the only new taxa in the volume.

Megalosaurus? "cachuensis" Weishampel, Barrett, Coria, Le Loeuff, Xu, Zhao, Sahni, Gomani and Noto, 2004
Middle Jurassic
Dapuka Group, Xinjiang, China
Comments- This is only present as a name in the Dinosaur Distribution chapter, so is a nomen nudum. It may be the same as the nomen nudum Megalosaurus? "dapukensis" Zhao 1985, which is not in the book.


"Aniksosaurus" Martinez vide anonymous, 1997
"A. darwini" Weishampel, Barrett, Coria, Le Loeuff, Xu, Zhao, Sahni, Gomani and Noto, 2004
Cenomanian-Coniacian, Late Cretaceous
Bajo Barreal Formation, Argentina
Comments- The name "Aniksosaurus darwini" has been on the internet since 2001, but this is its first publication in print. It is merely a name in the Dinosaur Distribution chapter though, so is still a nomen nudum.


Rinchenia Osmolska, Currie and Barsbold, 2004
R. mongoliensis (Barsbold, 1986) Osmolska, Currie and Barsbold, 2004
= Oviraptor mongoliensis Barsbold 1986
Early Maastrichtian, Late Cretaceous
Nemegt Formation, Mongolia
Comments. Osmolska et al. attribute this name to Barsbold 1997, but the latter use was as a nomen nudum. I don't see anything that would bar Osmolska et al. from having the name attributed to them for this chapter though. It's diagnosed, illustrated, and Barsbold 1986 and Oviraptor mongoliensis are explicitly stated as being the species and reference it's based on.


Now, what theropods (and basal saurischians) aren't included in The Dinosauria? Well, most nomina nuda aren't, but there are exceptions. The following nomina nuda are in the book, though italicized and not surrounded by quotation marks-
*Megalosaurus? "cachuensis"
*"Ngexisaurus dapukanensis"
Szechuanosaurus "yandonensis" (Basal Tetanurae)
Tyrannosaurus "lanpingensis" (Tyrannosauroidea)
*"Aniksosaurus darwini"
*"Proornis coreae"
"Paleopteryx thomsoni" (Dromaeosauridae)
"Holbotia ponomarenkoi" (Basal Avialae)
The four marked by asterisks are only in the Dinosaur Distribution chapter.


Moving to valid taxa, the following are only mentioned in the text, not in an informational table.
Shuvosaurus (Ornithomimosauria)
Dilophosaurus? sinensis (Ceratosauria)
Longipteryx (Basal Avialae)
Shuvosaurus is considered non-dinosaurian (though Rauhut's 1997 opinion is mentioned; his 2000/2003 one is not), and the other two also have Dinosaur Distribution entries. Longipteryx is only mentioned in the context of having feathers, but there are a few sentences about D? sinensis.


Calamosaurus is never mentioned, as it is called Calamospondylus foxi in the Basal Tetanurae table.

The Ceratosauria table fails to differentiate Longosaurus from Coelophysis? longicollis, though they have different holotypes.

The following are present as their original species, but their new genera are not listed-
Spinostropheus (as Elaphrosaurus gautieri) (Ornithomimosauria) 2004
Explorornis (as Enantiornis walkeri) (Basal Avialae) 1998
Incolornis (as Enantiornis martini) (Basal Avialae) 1998
Guildavis (as Ichthyornis tener) (Basal Avialae) 2004


There are many taxa which have Dinosaur Distribution entries, but are otherwise not listed. Publication dates are listed below if they are recent.
Arctosaurus
Macrodontophion
"Megalosaurus" obtusus
"Megalosaurus" cloacinus
"Plateosaurus" elizae
Sinosaurus
Zupaysaurus 2003
Compsosuchus
Rajasaurus 2003
"Massospondylus" rawesi
Megalosaurus? mersensis
"Morosaurus" marchei
Lukousaurus
Aviatyrannis 2003
Huaxiagnathus 2004
Praeornis
Yixianosaurus 2003
Dromaeosauroides 2003
Shenzhouraptor 2002
Jeholornis 2002
"Cathayornis" caudatus 1997
Cuspirostrisornis 1997
Largirostrornis 1997
Longchengornis 1997
Liaoxiornis 1999
Longirostravis 2003
Yanornis 2001
Yixianornis 2001
Songlingornis 1997
Eurolimnornis
Palaeocursornis
Hesperornis rossicus 1993
Some of these were probably too new, and others seem to lack a proper chapter (possible Dinosauria). A few are poorly known, but I can't help but think someone goofed in not including Compsosuchus, and that Padian could have done a better job on the Basal Avialae chapter (Eocathayornis, Halimornis, Sapeornis and Epidendrosaurus are included, all from 2002). Strangely, Confuciusornis chuonzhous and C. suniae are included, but not the enantiornithine taxa from that same volume.


In general, neornithines aren't treated by the book. The only one listed in a table is "Graculavus" lentus (which is now Austinornis). However, about half the known Mesozoic neornithines have Dinosaur Distribution entries. I can't figure out the reason some don't, since some of the best known taxa (e.g. Cimolopteryx, Graculavus) are missing. In any case, the following are absent from the book-
Ceramornis
Gallornis
Gobipipus
"Palaeotringa" vetus
Cimolopteryx
Graculavus
"Lonchodytes" priscus
Torotix
Lonchodytes


Finally, the following taxa are not found anywhere in the book at all. Again, recent taxa have publication dates listed.
Picrodon
"Ornithocheirus" hilsensis
Pycnonemosaurus 2002
Rugops 2004
Suchosaurus
Dilong 2004
Tyrannosaurus? zhuchengensis 2002
Mirischia 2004
Asiamericana 1995
Mei 2004
Sinusonasus 2004
Graciliraptor 2004
Cryptovolans 2002
Atrociraptor 2004
Jixiangornis 2002
Yandangornis 1999
Omnivoropteryx 2002
Jinzhouornis yixianensis 2002
Jinzhouornis zhangjiyingia 2002
Abavornis 1998
Catenoleimus 1998
Piksi 2002
Aberratiodontus 2004
"Cathayornis" aberransis 2002
Vescornis 2004
Explorornis? walkeri 1998
Canadaga 1999
Hesperornis bairdi 2002
Hesperornis chowi 2002
Hesperornis macdonaldi 2002
Hesperornis mengeli 2002
Parascaniornis
Iaceornis 2004


Again, Picrodon and Asiamericana might be deemed possibly dinosaurian, while Suchosaurus and "Ornithoscheirus" hilsensis are poorly known. It's odd for Microraptor gui to be included, but not the earlier named Cryptovolans. Yandangornis and Panteleev's Soviet bird paper seem to have slipped by Padian.

Overall, I'm impressed with the number of recently described taxa in the book, especially with Weishampel et al.'s job in the Dinosaur Distributions chapter.

Mickey Mortimer