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Centrosaurus brinkmani, new ceratopsian
Ryan, M.R. and Russell, A.P. (2005). A new centrosaurine ceratopsid from
Formation of Alberta and its implications for centrosaurine taxonomy and
systematics. Can. J. Earth Sci. 42: 1369?1387.
Abstract: "_Centrosaurus brinkmani_ (sp. nov) is distinguished from
_Centrosaurus apertus_ by key features of its cranial ornamentation,
including the shape and orientation of the postorbital horn and parietal
ornamentation at parietal locus 3, the shape of the parietal ornamentation
at locus 2, and the possession of accessory parietal ossifications developed
as short spines on the caudal parietal ramus. This species is restricted to
the Oldman Formation of southern Alberta and is the oldest ceratopsid
represented by diagnostic material in Canada. Phylogenetic analysis of the
Centrosaurinae suggests that the development of spike-like ornamentation at
the parietal locus 3 parietal locus is inversely related to the development
of the P1 parietal ornamentation."