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Re: pterosaur/bat wing membranes
... Q. was uniquely capable of going where no pterosaur before was
capable: to the deeper parts of the pond. And you can see this
phylogenetically with ever increasing size, following an initial bump
in relative neck length, metacarpal length and distal wing shrinkage
in smaller Q-ish sisters. Poling (hand only) pterosaur ichnites of
smaller genera show that at least some pterosaurs were doing this.
Phylogeny also supports this hypothesis in that Quetzalcoatlus, was
descended from Dorygnathus, which, with its relatively short wingspan
and a wide-mouth raking dentition, appears to have been a
sand/mud-sifter, perhaps seeking crustaceans...
The only thing about habitual "poling" that worries me is that the
animal cannot take off while wading in that manner. If that was a
major mode of feeding, it would leave a Quetz. individual very
vulnerable much of the time.