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Coelurosaur Analysis Update featuring Juravenator, Guanlong, Hagryphus, etc.



Amazing that I haven't done one of these since October - http://dml.cmnh.org/2005Oct/msg00355.html.

- added Guanlong, Hagryphus, Unenlagia paynemili, IGM 100/972 and 100/974 (Velociraptor/troodontid hatchling skulls described by Norell et al., 1994), WDC DML 001 (undescribed Morrison troodontid - preliminary codings), CAGS-IG-02-0901 (enantiornithine wing described by You et al., 2005), Hongshanornis and "Archaeorhychus".
- split Caenagnathidae into Caenagnathus holotype, Chirostenotes holotype, ROM 43250 (Chirostenotes described by Sues, 1997), RTMP 79.20.1 (Chirostenotes described by Currie and Russell, 1988), the undescribed CMNH/Triebold caenagnathid, and Elmisaurus rarus.
- updated codings for Acrocanthosaurus, Dilong, Juravenator (formerly Borsti), Yanornis and Yixianornis.
- all qualitative cranial characters coded for non-pygostylian birds, dromaeosaurids and troodontids.


90% majority rule consensus for the first 50000 most parsimonious trees-

|--Monolophosaurus
`--+--+--Acrocanthosaurus
  |  `--+--Allosaurus
  |     `--+--Sinraptor
  |        `--Megaraptor
  `--+--+--Fukuiraptor
     |  |--Stokesosaurus
     |  `--+--Itemirus
     |     `--"Alashansaurus"
     |*-Iliosuchus
     `--+--+--Dryptosaurus
        |  `--+--Tanycolagreus
        |     `--Coelurus
        `--+--+--Labocania
           |  `--+--+--Richardoestesia
           |     |  `--+--Guanlong
           |     |     `--Dilong
           |     `--+--+--Eotyrannus
           |        |  `--SMNS 58023
           |        `--+--Appalachiosaurus
           |           `--+--Albertosaurus
           |              `--+--Gorgosaurus
           |                 `--+--Daspletosaurus
           |                    `--+--Tarbosaurus
           |                       `--+--Bagaraatan
           |                          `--+--Aviatyrannis
           |                             `--Tyrannosaurus
           |*--Calamosaurus
           `--+--+--Nqwebasaurus
              |  `--+--Mirischia
              |     `--Ornitholestes
              `--+--NGMC 2124
                 `--+--Proceratosaurus
                    `--+--+--Juravenator
                       |  `--+--Huaxiagnathus
                       |     `--Sinosauropteryx
                       `--+--Scipionyx
                          `--+--Compsognathus
                             `--+--Archaeornithomimus
                                `--+--+--"Grusimimus"
                                   |  `--+--Pelecanimimus
                                   |     `--+--+--Garudimimus
                                   |        |  `--+--Sinornithomimus
                                   |        |     `--Ornithomimidae
                                   |        `--+--ROM 43250
                                   |           `--+--Anserimimus
                                   |              `--+--YPM 1996
                                   |                 `--+--Harpymimus
                                   |                    `--Shenzhousaurus
                                   `--Maniraptora

|--+--Patagonykus
|  `--+--Alvarezsaurus
|     `--+--Parvicursor
|        `--+--Shuvuuia
|           `--Mononykus
`--+--Therizinosaurus
  `--+--+--Alxasaurus
     |  |--Nothronychus
     |  |--Erliansaurus
     |  `--+--+--Beipiaosaurus
     |     |  `--Neimongosaurus
     |     `--+--Nanshiungosaurus
     |        `--+--Segnosaurus
     |           `--Erlikosaurus
     `--+--Caudipteryx
        `--+--Protarchaeopteryx
           `--+--+--+--Falcarius
              |  |  `--Incisivosaurus
              |  `--+--Caenagnathus
              |     |--Chirostenotes
              |     |--CMNH caenagnathid
              |     |--Hagryphus
              |     `--+--Microvenator
              |        `--+--ZPAL MgD-I/95
              |           `--+--+--Nemegtomaia
              |              |  `--Heyuannia
              |              `--+--Nomingia
              |                 `--+--+--Rinchenia
              |                    |  `--NXMV
              |                    `--+--Ingenia
              |                       `--+--+--Deinocheirus
              |                          |  `--+--Shixinggia
              |                          |     `--Citipati
              |                          `--+--+--Conchoraptor
              |                             |  `--Khaan
              |                             `--+--Elmisaurus
              |                                `--Oviraptor
              `--+--Avimimus
                 `--Eumaniraptora

|--+--Archaeornithoides
|  `--+--+--Unenlagia comahuensis
|     |  `--+--+--Yandangornis
|     |     |  `--Shenzhouraptor
|     |     `--+--+--Microraptor gui
|     |        |  `--+--Microraptor zhaoianus
|     |        |     `--Cryptovolans
|     |        `--+--NGMC 91
|     |           `--+--Graciliraptor
|     |              `--+--+--IGM 100/1015
|     |                 |  `--+--Bambiraptor
|     |                 |     `--+--Sinornithosaurus
|     |                 |        `--RTMP 79.20.1
|     |                 `--+--+--Saurornitholestes
|     |                    |  |*-Variraptor
|     |                    |  `--+--Deinonychus
|     |                    |     `--+--Achillobator
|     |                    |        `--Dromaeosaurus
|     |                    |*-Pyroraptor
|     |                    |*-Unenlagia paynemili
|     |                    `--+--Atrociraptor
|     |                       `--+--Velociraptor
|     |                          `--+--Utahraptor
|     |                             `--Adasaurus
|     `--+--+--Scansoriopterygidae
|        |  `--+--IGM 100/974
|        |     `--+--Mei
|        |        `--+--Archaeopteryx
|        |           `--Wellnhoferia
|        `--+--+--Rahonavis
|           |  `--Buitreraptor
|           `--+--+--Ukhaa Tolgod troodontid
|              |  `--+--Sinovenator
|              |     `--+--Jinfengopteryx
|              |        `--Yixianosaurus
|              `--+--WDC DML 001
|                 `--+--Sinusonasus
|                    `--+--+--IGM 100/44
|                       |  `--+--IGM 100/972
|                       |     `--Byronosaurus
|                       `--+--Sinornithoides
|                          `--+--Troodon
|                             `--+--Saurornithoides junior
|                                `--Saurornithoides mongoliensis
`--+--+--Sapeornis
  |  `--Omnivoropteryx
  `--+--+--Confuciusornis
     |  `--Changchengornis
     `--+--Jibeinia
        `--+--Aberratiodontus
           `--+--+--Protopteryx
              |  `--+--CAGS-IG-02-0901
              |     `--+--+--Vescornis
              |        |  `--+--Eoalulavis
              |        |     `--Liaoningornis
              |        `--+--Sinornis
              |           `--+--Eocathayornis
              |              `--+--Jixiangornis
              |                 `--+--Longipteryx
              |                    `--Longirostravis
              `--+--Iberomesornis
                 `--+--+--Eoenantiornis
                    |  `--LP-4450-IEI
                    `--+--Concornis
                       `--+--+--Gobipteryx
                          |  `--Boluochia
                          `--+--+--Songlingornis
                             |  `--+--Patagopteryx
                             |     `--Ambiortus
                             `--+--Yanornis
                                `--+--Yixianornis
                                   `--+--Hesperornithes
                                      `--+--+--Ichthyornis
                                         |  `--IVPP V11309
                                         `--+--Iaceornis
                                            `--+--Apsaravis
                                               `--+--+--Hongshanornis
                                                  |  `--"Archaeorhychus"
                                                  `--+--Noguerornis
                                                     `--+--Neuquenornis
                                                        `--Enantiornis

Note Guanlong is sister to Dilong, not basal to it. Juravenator comes out sister to Huaxiagnathus and Sinosauropteryx, but not in Compsognathidae. Hagryphus clusters with most caenagnathids at the base of Caenagnathoidea, but Elmisaurus is an oviraptorid, ROM 43250 is an ornithomimosaur and RTMP 79.20.1 is a dromaeosaurid. Apparently caenagnathids need more characters to form a clade. Unenlagia paynemili has an uncertain position, but it's never close to U. comahuensis. One of the hatchling skulls is sister to Byronosaurus, but the other is by Mei and archaeopterygids. I suspect it may be from the Ukhaa Tolgod basal troodontid (called Shuvuuia on the AMNH website). The fact a hatchling nested sister to its supposed adult form is encouraging when it comes to taxa like Scipionyx and Juravenator, especially considering that I haven't adjusted codings for ontogeny yet. Buitreraptor and Rahonavis are still closer to troodontids and archaeopterygids than to dromaeosaurids.

"Improvements" since last time-
- Megaraptor is an allosauroid.
- Labocania is a tyrannosauroid.
- Appalachiosaurus is outside Tyrannosauridae.
- Juravenator isn't an arctometatarsalian.
- Archaeornithoides isn't an ornithomimosaur.
- Erliansaurus and Nothronychus are therizinosaurs.
- Therizinosaurus is closer to Therizinosauria.
- Microvenator is basal to other oviraptorids.
- Jinfengopteryx is a troodontid.
- Sapeornis and Omnivoropteryx aren't oviraptorosaurs.
- Apsaravis and Iaceornis are closer to carinates than yanornithids.

Things that have worsened since last time-
- Stokesosaurus isn't a tyrannosauroid.
- Gorgosaurus isn't an albertosaurine.
- Archaeornithomimus isn't an ornithomimosaur.
- Avimimus, Caudipteryx and Protarchaeopteryx aren't oviraptorosaurs.
- Falcarius isn't a therizinosaur.
- Jixiangornis is a pygostylian.
- Noguerornis, Neuquenornis and Enantiornis are deep within Ornithuromorpha.

Mickey Mortimer