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Birds have "thumb" genes like alligators: free pdf

From: Ben Creisler

In case this item has not been mentioned, a new study of 
alligator digits adds weight to theory that a "hometic 
frameshift" occurred in birds, so that wing fingers one, 
two and three shifted to the embryological positions of 
fingers two, three and four.

Vargas AO, Kohlsdorf T, Fallon JF, VandenBrooks J, Wagner 
GP (2008) The Evolution of HoxD-11 Expression in the Bird 
Wing: Insights from Alligator mississippiensis. PLoS ONE 3
(10): e3325
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