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Re: Citing combinations with subgenera
The ICZN states: 'If it is desired to cite both the author of a
species-group nominal taxon and the person who first transferred it to
another genus, the name of the person forming the new combination should
follow the parentheses that enclose the name of the author of the
and Pteranodon (Geosternbergia) sternbergi (Harksen, 1966) Miller, 1978 is
But this is exactly what your quote of the ICZN says -- except that that
applies to genera rather than subgenera... but maybe you should use it
anyway. After all, the species name is due to Harksen, and the name as a
whole in that form is due to Miller.
Mike Keesey suggested Pteranodon (Geosternbergia) sternbergi Harksen, 1966
per Miller, 1978, but this seems odd to me.
Among other things because _per_ isn't mentioned in the ICZN. (That also
means it can't be wrong, though.)