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New Darwinopterus species (Pterosauria)

From: Ben Creisler

LÜ, J., XU, L., CHANG, H. and ZHANG, X. (2011).
A New Darwinopterid Pterosaur from the Middle Jurassic of Western Liaoning,
Northeastern China and its Ecological Implications. 
Acta Geologica Sinica - English Edition 85(3): 507?514, June 2011
doi: 10.1111/j.1755-6724.2011.00444.x

A new species of Darwinopterus, D. robustodens sp. nov. is described and
named. Based on the new specimen, the diagnostic characters of
Darwinopterus are amended and include: rostral dentition composed of
well-spaced, spike-like teeth; the longest teeth are confined to the
anterior half of the tooth row; tooth alveoli have raised margins;
nasoantorbital fenestra confluent; inclined quadrate; elongate cervical
vertebrae with low neural spine and reduced or absent ribs; long tail of
more than 20 caudals partially enclosed by filiform extensions of the pre-
and postzygapophyses; short metacarpus less than 60 per cent length of
humerus, fifth toe with two elongate phalanges and curved second pedal
phalanx of the fifth toe with the angle between the proximal and distal
segments about 130 degrees. The complete specimen of the new pterosaur D.
robustodens sp. nov. provides much more osteological information. The
differences in tooth morphologies between Darwinopterus modularis and D.
robustodens sp. nov. suggest that they filled different ecological niches.
The hard integument-bearing Coleoptera may have been the main food source
of Darwinopterus robustodens.

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