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Crocodilians by the millions! Special issue of Zoological Journal!

From: Ben Creisler

This item came out yesterday but has not been mentioned yet on the
DML. A special supplement issue of the Zoological Journal of the
Linnean Society is devoted to the 1st Symposium on the Evolution of
Crocodyliforms. All the pdfs are FREE!


Many Mesozoic crocs are discussed and some new genera and species are
described. I'll cite the new taxa for now. The correct date for these
taxa is a bit murky--it's officially the December 2011 issue but it
was put on line (published?) in 2012. The "how to cite" feature says


KELLNER, A. W. A., CAMPOS, D. A., RIFF, D. and DE ANDRADE, M. B. (2011)
A new crocodylomorph (Sphagesauridae, Notosuchia) with horn-like
tubercles from Brazil. Zoological Journal of the Linnean Society, 163:
doi: 10.1111/j.1096-3642.2011.00712.x

A new species of a bizarre notosuchian mesoeucrocodylian is reported
here. Caryonosuchus pricei gen. et sp. nov. was found in the outcrops
of the Adamantina Formation (Campanian-Masstrichtian) in São Paulo
State, Brazil, and shows a typical sphagesaurid dentition: strong and
short teeth, obliquely implanted with the crown of the upper teeth
showing a rounded anteriolabial margin and a strong compressed
posteriolingual edge developed into a carina, ornamented by developed
ridges and denticles. Amongst the diagnostic features of the new taxon
are the presence of horn-like tubercles on the premaxilla and maxilla,
never reported in this group before. The occurrence of C. pricei
increases the diversity of sphagesaurids and confirms that all members
of this clade, only recorded in Late Cretaceous deposits from Brazil
so far, share the same dentition.


A new Berriasian species of Goniopholis (Mesoeucrocodylia, Neosuchia)
from England, and a review of the genus.
Zoological Journal of the Linnean Society, 163: S66-S108.
doi: 10.1111/j.1096-3642.2011.00709.x

A new species, Goniopholis kiplingi sp. nov., based on an
exceptionally preserved skull from the Lower Cretaceous of England is
described in detail. It shows great similarity with Goniopholis simus
and Goniopholis baryglyphaeus, but can be distinguished by the
presence of longer lachrymals, smooth (not edged) dorsal surface of
the quadrate, and proportionally longer rostrum. A comprehensive
phylogenetic analysis of Mesoeucrocodylia (104 taxa; 486 characters)
focused on goniopholidids (14 species) places G. kiplingi as
sister-group of G. simus, and as part of a monophyletic group also
containing G. baryglyphaeus. The relationships of Nannosuchus
gracilidens and three undescribed European taxa are explored, and
preliminary analyses of Denazinosuchus kirtlandicus (Upper Cretaceous,
USA) and 'Goniopholis' phuwiangensis (Lower Cretaceous, Thailand) are
presented. The assignment of taxa to the genus Goniopholis is
discussed. Goniopholis, in its traditional sense, is considered
paraphyletic and a restricted updated definition is proposed, with
comments on the evolution of other goniopholidids. Morphological
characteristics of fragmentary material attributed to Goniopholis are
not considered sufficient to secure their generic/specific assignment,
and provide no support for the presence of Goniopholis in Gondwanan
and/or Upper Cretaceous sedimentary units. Currently Goniopholis is
restricted to the Upper Jurassic-Lower Cretaceous of Europe.



KELLNER, A. W. A., FIGUEIREDO, R. G., AZEVEDO, S. A. K. and CAMPOS, D. A. (2011)
A new cretaceous notosuchian (Mesoeucrocodylia) with bizarre dentition
from Brazil. Zoological Journal of the Linnean Society, 163:
doi: 10.1111/j.1096-3642.2011.00711.x

A new species of Notosuchia, Labidiosuchus amicum gen. et sp. nov., is
described based on an incomplete lower jaw (DGM 1480-R) from the Upper
Cretaceous Marília Formation (Maastrichtian) recovered from a quarry
near the Peirópolis municipality, Minas Gerais State, Southeastern
Brazil. The mandibular symphysis is long, strong anterodorsally
projected and 'Y-shaped'. The bizarre dentition is formed by at least
eight teeth placed in a symphyseal tooth battery, some located lateral
to each other. The first pair is larger than all others and
procumbent. Some teeth are obliquely implanted (anterolabially to
posterolingually) and have sub circular to elliptical outline. At
least the posterior teeth are single cuspidate with acute apex.
Labidiosuchus amicum shows a rather bizarre dentition, increasing the
taxonomic diversity and potential feeding strategies of notosuchian



CLARK, J. M. (2011)
A new shartegosuchid crocodyliform from the Upper Jurassic Morrison
Formation of western Colorado.
Zoological Journal of the Linnean Society, 163: S152-S172.
doi: 10.1111/j.1096-3642.2011.00719.x

A small new basal crocodyliform, Fruitachampsa callisoni gen. nov.,
sp. nov., is represented by several partial skeletons from the
Morrison Formation at the Fruita Paleontological Area near Grand
Junction, Colorado. It is placed in the Family Shartegosuchidae
Efimov, 1988, previously comprising three genera from the Late
Jurassic locality of Shar Teeg in western Mongolia and possibly a
fourth genus from the Early Cretaceous of Siberia. Shartegosuchids
share a sculpted palatal surface of the pterygoids, the absence of a
mandibular fenestra, and posterior maxillary teeth and post-caniniform
dentary teeth with a flat and horizontal apical region and vertical
crenulations extending basally from it. Fruitachampsa and
Shartegosuchus form a clade supported by ventral half of the lacrimal
tapering ventroposteriorly, sculpturing on palatines, and lower teeth
absent anterior to caniniforms. The shartegosuchids are most
parsimoniously considered to be outside of the mesoeucrocodylian clade
and are possibly allied with the Asian taxa Shantungosuchus,
Sichuanosuchus, and Zosuchus. Fruitachampsa is unusual in possessing a
series of small protuberances along the occipital margin of the
parietal and squamosal and procoelous vertebrae, and lacking an
antorbital fenestra or fossa. This is the first occurrence of a
shartegosuchid in North America, and the close relationship of
Fruitachampsa with Shartegosuchus nested among other Asian taxa
indicates it dispersed to North America from Asia.



Early eusuchia crocodylomorpha from the vertebrate-rich Plattenkalk of
Pietraroia (Lower Albian, southern Apennines, Italy).
Zoological Journal of the Linnean Society, 163: S199-S227.
doi: 10.1111/j.1096-3642.2011.00718.x

The locality of Pietraroia (Lower Albian, southern Apennines, Italy)
has provided two fully articulated crocodylomorphs, exposed in ventral
and in dorsal aspect, which are described here as representing a new
species of a new genus, Pietraroiasuchus ormezzanoi gen. nov, sp. nov.
The new taxon is found to be the sister taxon of Pachycheilosuchus
trinquei from the Albian of the Glen Rose Formation, Texas.
Pietraroiasuchus ormezzanoi resolves the phylogenetic position of the
controversial P. trinquei, and is crucial in enabling an extensive
understanding of the family Hylaeochampsidae. Phylogenetic analysis
places Hylaeochampsa vectiana as a sister group of Iharkutosuchus
makadii plus Pachycheilosuchus and Pietraroiasuchus. The phylogenetic
result reveals the presence of an evolutionary mosaicism within
non-crocodylian eusuchians. Mosaic features in Pietraroiasuchus are
denoted by the combination of primitive character states, such as the
position of the choana with its anterior margin formed by an inverted
V-shaped palatine processeses, and the presumed presence of a tiny
antorbital fossa, in conjunction with derived states involving the
slight vertebral procoelia, the presence of tetraserial segmented
dermal armour with an accessory lateral row, and isolated nuchal
osteoderms. The disjoint occurrence between Pachycheilosuchus and
Pietraroiasuchus species suggests that Pietraroia was a refuge island
inhabited by endemic forms.


Not a new taxon, but relevant to dinosaurs:

RIFF, D. and KELLNER, A. W. A. (2011)
Baurusuchid crocodyliforms as theropod mimics: clues from the skull
and appendicular morphology of Stratiotosuchus maxhechti (Upper
Cretaceous of Brazil).
Zoological Journal of the Linnean Society, 163: S37-S56.
doi: 10.1111/j.1096-3642.2011.00713.x

The Baurusuchidae crocodyliforms are usually interpreted as active
terrestrial predators, but only some positive evidence of such habits
has been described to date, mainly the relative position of external
nares and orbits. Here we describe features that support this view in
a complete specimen of the Baurusuchidae Stratiotosuchus maxhechti,
and have executed a parsimony analysis to confirm their phylogenetic
position. S. maxhechti exhibits theropodomorph features that have been
previously recognized in skulls of the Baurusuchidae, as well as
postcranial characteristics related to a parasagittal gait, showing
that the similarities between the Baurusuchidae and theropods extend
beyond the cranial morphology. These include a well-developed
supracetabular crest, a relatively medially offset femoral head and a
caudally orientated calcaneal tuber. The orientations of the surfaces
for muscular attachments imply that the appendicular movements of S.
maxhechti were mainly anteroposterior, with abduction significantly
constrained. S. maxhechti presents features that mimic some present in
theropods, including a 'fossa brevis' on the ilium and tubercles on
the ischium and femur similar to the obturator process and accessory
trochanter. The relative proportions of the femur, tibia, and longer
metatarsal are more similar to those of Postosuchus than to other
Crocodylomorpha. In the skull, besides the theropodomorph (ziphodont)
dentition concentrated in the anterior half of the rostrum, the
baurusuchids are remarkable by the fusion of the nasals, which can be
related to a large resistance against feeding forces acting on a
high-profile skull. The appendicular morphology of S. maxhechti
strengthens the interpretation that the Baurusuchidae were active
land-dwelling predators in the Upper Cretaceous of south-eastern
Brazil, occuping ecological niches typical of small to medium-sized
theropod dinosaurs.