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Nasutoceratops, centrosaurine ceratopsid from Campanian of Utah (free pdf)

From: Ben Creisler

The paper for Nasutoceratops is now officially out and in open
access.[I'm glad the spelling was revised from the cut-and-paste
"Nasutuceratops" to use the standard Neo-Latin "o" as a combining

Scott D. Sampson, Eric K. Lund, Mark A. Loewen, Andrew A. Farke and
Katherine E. Clayton (2013)
A remarkable short-snouted horned dinosaur from the Late Cretaceous
(late Campanian) of southern Laramidia.
Proceedings of the Royal Society B 280 No. 1766 20131186
doi: 10.1098/rspb.2013.1186

Free pdf:

The fossil record of centrosaurine ceratopsids is largely restricted
to the northern region of western North America (Alberta, Montana and
Alaska). Exceptions consist of single taxa from Utah (Diabloceratops)
and China (Sinoceratops), plus otherwise fragmentary remains from the
southern Western Interior of North America. Here, we describe a
remarkable new taxon, Nasutoceratops titusi n. gen. et sp., from the
late Campanian Kaiparowits Formation of Utah, represented by multiple
specimens, including a nearly complete skull and partial postcranial
skeleton. Autapomorphies include an enlarged narial region, pneumatic
nasal ornamentation, abbreviated snout and elongate, rostrolaterally
directed supraorbital horncores. The subrectangular parietosquamosal
frill is relatively unadorned and broadest in the mid-region. A
phylogenetic analysis indicates that Nasutoceratops is the sister
taxon to Avaceratops, and that a previously unknown subclade of
centrosaurines branched off early in the group's history and persisted
for several million years during the late Campanian. As the first
well-represented southern centrosaurine comparable in age to the bulk
of northern forms, Nasutoceratops provides strong support for the
provincialism hypothesis, which posits that Laramidia—the western
landmass formed by inundation of the central region of North America
by the Western Interior Seaway—hosted at least two coeval dinosaur
communities for over a million years of late Campanian time.