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Ornithocheirus (Pterosauria) taxonomy (Lonchodraco, Cimiliopterus and Campsipterus new genera) (free pdf)

From: Ben Creisler

A new monograph in open-access Zookeys:

Taissa Rodrigues &  Alexander Kellner (2013)
Taxonomic review of the Ornithocheirus complex (Pterosauria) from the
Cretaceous of England.
ZooKeys 308: 1–112,
doi: 10.3897/zookeys.308.5559

Over a decade after the last major review of the Cambridge Greensand
pterosaurs, their systematics remains one of the most disputed points
in pterosaur taxonomy. Ornithocheiridae is still a wastebasket for
fragmentary taxa, and some nomenclatural issues are still a problem.
Here, the species from the Cretaceous of England that, at some point,
were referred in Ornithocheirus, are reviewed. Investigation of the
primary literature confirmed that Criorhynchus should be considered an
objective junior synonym of Ornithocheirus. Taxonomic review of more
than 30 species known from fragmentary remains showed that 16 of them
are undiagnosable (nomina dubia): Palaeornis cliftii, Cimoliornis
diomedeus, Pterodactylus compressirostris, Pterodactylus fittoni,
Pterodactylus woodwardi, Ornithocheirus brachyrhinus, Ornithocheirus
carteri, Ornithocheirus crassidens, Ornithocheirus dentatus,
Ornithocheirus enchorhynchus, Ornithocheirus eurygnathus,
Ornithocheirus oxyrhinus, Ornithocheirus scaphorhynchus,
Ornithocheirus tenuirostris, Ornithocheirus xyphorhynchus, and
Pterodactylus sagittirostris. Fourteen species are considered valid,
and diagnoses are provided to all of them: Ornithocheirus simus,
Lonchodraco giganteus comb. n., Lonchodraco machaerorhynchus comb. n.,
Lonchodraco(?) microdon comb. n., Coloborhynchus clavirostris,
‘Ornithocheirus’ capito, Camposipterus nasutus comb. n.,
Camposipterus(?) sedgwickii comb. n., Camposipterus(?) colorhinus
comb. n., Cimoliopterus cuvieri comb. n., ‘Ornithocheirus’ polyodon,
‘Ornithocheirus’ platystomus, ‘Pterodactylus’ daviesii, and
‘Ornithocheirus’ denticulatus. These species are referred in the
genera Ornithocheirus, Lonchodraco gen. n., Coloborhynchus,
Cimoliopterus gen. n., and Camposipterus gen. n., but additional
genera are probably present, as indicated by the use of single
quotation marks throughout the text. A cladistic analysis demonstrates
that Anhangueridae lies within a newly recognized clade, here named
Anhangueria, which also includes the genera Cearadactylus,
Brasileodactylus, Ludodactylus, and Camposipterus. The anhanguerian
‘Cearadactylus’ ligabuei belongs to a different genus than
Cearadactylus atrox. Lonchodraconidae fam. n. (more or less equivalent
to Lonchodectidae sensu Unwin 2001) is a monophyletic entity, but its
exact phylogenetic position remains uncertain, as is the case of
Ornithocheirus simus. Therefore, it is proposed that Ornithocheiridae
should be constricted to its type species and thus is redundant. Other
taxa previously referred as “ornithocheirids” are discussed in light
of the revised taxonomy.

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