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Dinosaurs in special issue of Journal of Iberian Geology (free pdfs)

Ben Creisler

The new issue of Journal of Iberian Geology 41(10) 2015 Dinosaur
Palaeontology and Environment is now online in open access:



F. Torcida Fernández-Baldor, X. Pereda-Suberbiola & D.B. Weishampel (2015)
Dinosaur Palaeontology and Environment.
Journal of Iberian Geology 41(1): 7-10
doi: http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48650


A. Pérez-García (2015)
Revision of the British record of Tropidemys (Testudines,
Plesiochelyidae) and recognition of its presence in the Late Jurassic
of Portugal.
Journal of Iberian Geology 41(1): 11-20
doi: http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48651

The record of coastal marine turtles belonging to Plesiochelyidae is
abundant in the Late Jurassic of Portugal. The material analyzed thus
far has been attributed to two taxa: Plesiochelys and Craspedochelys.
A specimen is presented here that allows extending the known diversity
of Portuguese Jurassic turtles. It is attributed to Tropidemys.
Although this taxon is relatively well known in the Kimmeridgian
record of Switzerland and Germany, no specific allocation performed
outside these countries can be, so far, confirmed. The detailed study
of the poorly known British taxon “Pelobatochelys” blakii allows its
specific validity to be confirmed here, being recognized as a member
of Tropidemys. The revision of this species and the analysis of the
new Portuguese specimen allow extending the knowledge regarding the
genus Tropidemys.


Jurassichelon, new taxon

A. Pérez-García (2015)
New data on the poorly-known Late Jurassic European turtles
Thalassemys and Enaliochelys and description of a new basal
eucryptodiran taxon.
Journal of Iberian Geology 41(1): 21-30
doi: http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48652

One of the most abundant clades of turtles in the Late Jurassic record
of Europe is Eucryptodira, represented by Plesiochelyidae,
Eurysternidae, Hylaeochelys and Thalassemys. The detailed study of a
British specimen preliminarily attributed to the plesiochelyid
Pelobatochelys” allows this attribution to be refuted. Rather, it is
recognized as the only confirmed specimen of the genus Thalassemys
outside Central Europe. The diversity of Eucryptodira in the Late
Jurassic of Europe is here increased, thanks to the revision of two
problematic specimens, not assignable to any of the four above
mentioned taxa. Enaliochelys chelonia, is a poorly-known British taxon
ignored since the late nineteenth century because of its putative
synonymy with Thalassemys hugii. The validity of this taxon is
supported here and a diagnosis for it is proposed for the first time.
The other specimen, from France (Oléron Island), was previously
attributed to “Thalassemys moseri”, which is currently considered
invalid. It is recognized as a representative of a new taxon,
Jurassichelon oleronensis gen. et sp. nov.

J. Parrilla-Bel & J.I. Canudo (2015)
Postcranial elements of Maledictosuchus riclaensis (Thalattosuchia)
from the Middle Jurassic of Spain.
Journal of Iberian Geology 41(1): 31-40
doi http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48653

Maledictosuchus riclaensis is a metriorhynchid crocodylomorph from the
Callovian (Middle Jurassic) of Ricla (Spain). It is the most basal
member of the Rhacheosaurini Tribe; it has recently been described and
defined by its cranial elements (an almost complete skull and part of
the lower jaw), but there were no data on the postcranial elements.
Associated with the skull three vertebrae were collected. These
vertebrae were preserved in black calcite nodules, and they have
recently been prepared. The postcranial elements of the
metriorhynchids are poorly documented, and usually badly preserved or
included in the matrix. Herein we describe the three vertebrae (part
of the holotype) of M. riclaensis. These comprise one cervical, one
dorsal and one caudal vertebra, which, like the skull, are well
preserved and lack postmortem distortion or deformation.

E. Puértolas-Pascual, J.I. Canudo & L.M. Sender (2015)
New material from a huge specimen of Anteophthalmosuchus cf. escuchae
(Goniopholididae) from the Albian of Andorra (Teruel, Spain):
Phylogenetic implications.
Journal of Iberian Geology 41(1): 41-56
doi: http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48654

In 2011 the partial skeleton of a goniopholidid crocodylomorph was
recovered in the ENDESA coal mine Mina Corta Barrabasa (Escucha
Formation, lower Albian), located in the municipality of Andorra
(Teruel, Spain). This new goniopholidid material is represented by
abundant postcranial and fragmentary cranial bones. The study of these
remains coincides with a recent description in 2013 of at least two
new species of goniopholidids in the palaeontological site of Mina
Santa María in Ariño (Teruel), also in the Escucha Formation. These
species are Anteophthalmosuchus escuchae, Hulkepholis plotos and an
undetermined goniopholidid, AR-1-3422. In the present paper, we
describe the postcranial and cranial bones of the goniopholidid from
Mina Corta Barrabasa and compare it with the species from Mina Santa
María. For the first time, we include the taxa from the Escucha
Formation in a phylogenetic analysis to establish their relationships
within Goniopholididae, adding to what is known of the goniopholidid
fossil assemblages from this time interval. The results indicate that
the specimen from Mina Corta Barrabasa is closely related to
Anteophthalmosuchus escuchae and may be the same species. Together
with Hulkepholis plotos and other taxa from England and Belgium, these
taxa form an endemic European clade. The Escucha Fm. goniopholidids
are the youngest Eurasian members of the group and may be the youngest
globally, depending on how North American taxa are eventually


J.M. Gasca, M. Moreno-Azanza, J.I. Ruiz-Omeñaca & J.I. Canudo (2015)
New material and phylogenetic position of the basal iguanodont
dinosaur Delapparentia turolensis from the Barremian (Early
Cretaceous) of Spain.
Journal of Iberian Geology 41(1): 57-70
doi: http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48655

Delapparentia turolensis Ruiz-Omeñaca, 2011 is the only iguanodont
taxon erected in the Barremian of Spain. It is described on the basis
of a partial postcranial skeleton discovered in the 1950s near the
village of Galve (Teruel Province), within the Camarillas Formation.
Recently, new remains from the same individual have been recovered,
and these are described here. Furthermore, after first-hand
examinations of the holotype, the phylogenetic position of this taxon
has been analysed for the first time, and its diagnosis is emended.
Delapparentia turolensis is a large-sized, basal iguanodont which
presents an autapomorphic, unusually high axial neural spine and a
unique combination of postcranial characters. The ilium morphology
differs from that of other basal iguanodonts and relates Delapparentia
to the Valanginian Barilium dawsoni from England, with whom it shares
two synapomorphies. In our phylogenetic analysis Delapparentia is
recovered in a polytomy with Kukufeldia, Lanzhousaurus, Barilium and
the clade equivalent to Iguanodontoidea.

J. Company, P. Cruzado-Caballero & J.I. Canudo (2015)
Presence of diminutive hadrosaurids (Dinosauria: Ornithopoda) in the
Maastrichtian of the south-central Pyrenees (Spain).
Journal of Iberian Geology 41(1): 71-81

In recent years a rich and diverse fauna of hadrosaurid dinosaurs has
been described in the Upper Cretaceous of the Pyrenees. Recent
fieldwork carried out in the upper Maastrichtian levels of the Tremp
Formation, in the south-central Pyrenees (province of Huesca,
northeastern Spain), has allowed us to recover diminutive fossil bones
referable to hadrosaurid dinosaurs. To date, small-sized specimens had
not been reported in the area. The remains consist of small vertebrae
and fragmentary long bones found in a relatively small area, so it is
assumed that they probably belong to individuals of a single
population. A morphological examination and a histological study
reveal that they represent specimens of advanced ontogenetic stage and
allow the identification of an undescribed taxon of small-bodied
hadrosaurids. In other parts of Europe, discoveries of small dinosaurs
have been linked to insularity. These findings bring to light the
smallest hadrosaurid known in Europe to date.


F. Ortega, N. Bardet , F. Barroso-Barcenilla, P.M. Callapez, O.
Cambra-Moo, V. Daviero-Gómez, V. Díez Díaz, L. Domingo, A. Elvira, F.
Escaso, M. García-Oliva, B. Gómez, A. Houssaye, F. Knoll, F.
Marcos-Fernández, M. Martín, P. Mocho, I. Narváez, A. Pérez-García, D.
Peyrot, M. Segura, H. Serrano, A. Torices, D. Vidal & J. L. Sanz
The biota of the Upper Cretaceous site of Lo Hueco (Cuenca, Spain).
Journal of Iberian Geology 41(1): 83-99
doi: http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48657

The Late Cretaceous (Campanian-Maastrichtian) fossil site of Lo Hueco
was recently discovered close to the village of Fuentes (Cuenca,
Spain) during the cutting of a little hill for installation of the
railway of the Madrid-Levante high-speed train. To date, it has
yielded a rich collection of well-preserved Cretaceous macrofossils,
including plants, invertebrates, and vertebrates. The recovered fossil
assemblage is mainly composed of plants, molluscs (bivalves and
gastropods), actinopterygians and teleosteans fishes, amphibians,
panpleurodiran (bothremydids) and pancryptodiran turtles, squamate
lizards, eusuchian crocodyliforms, rhabdodontid ornithopods, theropods
(mainly dromaeosaurids), and titanosaur sauropods. This assemblage was
deposited in a near-coast continental muddy floodplain crossed by
distributary sandy channels, exposed intermittently to brackish or
marine and freshwater flooding as well as to partial or total
desiccation events.

The Konzentrat-Lagerstatt of Lo Hueco constitutes a singular
accumulation of fossils representing individuals of some particular
lineages of continental tetrapods, especially titanosaurs, eusuchians
and bothremydid turtles. In the case of the titanosaurs, the site has
yielded multiple partial skeletons in anatomical connection or with a
low dispersion of their skeletal elements. A combination of new taxa,
new records of taxa previously known in the Iberian Peninsula, and
relatively common taxa in the European record compose the Lo Hueco
biota. The particular conditions of the fossil site of Lo Hueco and
the preliminary results indicate that the analysis of the geological
context, the floral and faunal content, and the taphonomical features
of the site provide elements that will be especially useful for
reassess the evolutionary history of some lineages of European Late
Cretaceous reptiles.


X. Pereda-Suberbiola, J.C. Corral, H. Astibia, A. Badiola, N. Bardet,
A. Berreteaga, E. Buffetaut, A.D. Buscalioni, H. Cappetta, L. Cavin,
V. Díez Díaz, E. Gheerbrant, X. Murelaga, F. Ortega, A. Pérez-García,
F. Poyato-Ariza, J.-C. Rage, J.L. Sanz & A. Torices (2015)
Late Cretaceous continental and marine vertebrate assemblages of the
Laño Quarry (Basque-Cantabrian Region, Iberian Peninsula): an update.
Journal of Iberian Geology 41(1): 101-124
doi: http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48658

The vertebrate-bearing beds of the Laño quarry (Condado de Treviño)
are among the most relevant sites from the Late Cretaceous of Europe.
Geologically, Laño and the adjacent region are set on the southern
limb of the South-Cantabrian Synclinorium (SE Basque-Cantabrian
Region, northern Iberian Peninsula). The Laño sites were discovered in
1984; thousands of bones and teeth, including microfossils, have been
collected during the prospection in the field and excavation
campaigns. The vertebrate remains occur at two different stratigraphic
horizons within a continental to shallow marine succession of Late
Campanian-Maastrichtian age. The lower horizon contains the Laño 1 and
Laño 2 sites, whereas the upper horizon contains the Albaina site. In
the Laño sites, three fossiliferous beds (called L1A, L1B and L2) are
known within an alluvial system composed mainly of fluvial sands and
silts. The sedimentary structures are consistent with channel areas
within an extensive braided river system. Based mainly on
stratigraphic correlations, the fluvial beds of Laño are regarded as
Late Campanian to Early Maastrichtian in age. These deposits have
yielded a very diverse vertebrate assemblage, which consists of nearly
40 species, including actinopterygians, lissamphibians, lepidosaurs,
turtles, crocodyliforms, dinosaurs, pterosaurs, and mammals. Seven
genera and ten species have been erected to date in Laño. With
reference to the marine vertebrate association of Albaina, it consists
of at least 37 species, including sharks and rays, actinopterygians,
mosasaurids, and plesiosaurs. Two genera and species of rhinobatoids
(family indet.) and two new species of rhinobatids have been erected
in Albaina. The fossil association indicates a Late (but not latest)
Maastrichtian age. Recently, isolated turtle and dinosaur fossils have
been discovered in the sublittoral beds of Albaina. The Laño quarry is
one of the most noteworthy Campanian-Maastrichtian vertebrate
localities of Europe by its taxonomic diversity, and provides useful
information about the composition and affinities of both continental
and marine vertebrate faunas from the latest Cretaceous of
southwestern Europe.


A. G. Sellés & B. Vila (2015)
Re-evaluation of the age of some dinosaur localities from the southern
Pyrenees by means of megaloolithid oospecies.
Journal of Iberian Geology 41(1): 125-139
doi: http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48659

Since the beginning of the 20th Century the Upper Cretaceous
continental and transitional marine deposits of the southern Pyrenees
have produced more than 220 dinosaur fossil localities. New
discoveries and advances in magnestostratigraphy and biostratigraphy
provide a robust biostratigraphical framework for the latest
Cretaceous dinosaur faunas in the region, although the age of some
localities has remained uncertain. Here, we re-evaluate the age of
some classic and new dinosaur fossil localities on the basis of the
potential dating of megaloolithid oospecies and provide parataxonomic
and age data for twenty-three localities with new megaloolithid egg
fossils. Further, we review the nomenclature and probable synonymies
of several of the most historically significant localities in the
southern Pyrenees. With the new age assignments proposed for some
significant localities (Basturs, Orcau-1, Els Nerets, Figuerola-2,
Suterranya-1), we claim that in the southern Pyrenees a) the
ankylosaurian dinosaurs survived beyond the early Maastrichtian-late
Maastrichtian boundary and coexisted with hadrosauroids; b) the
theropod record is scarce in the early Maastrichtian and the taxonomic
diversity (richness) of theropods is notably higher in the late
Maastrichtian; and c) the megaloolithid egg record assigned to
sauropods is continuous through the entire Maaastrichtian but is
scarce in the Upper Campanian.


F. Torcida Fernández-Baldor, I. Díaz-Martínez, R. Contreras, P.
Huerta, D. Montero & V. Urién (2015)
Unusual sauropod tracks in the Jurassic-Cretaceous interval of the
Cameros Basin (Burgos, Spain).
Journal of Iberian Geology 41(1): 141-154
doi: http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48660

Thee Las Sereas site includes at least 14 ichnological outcrops along
5.6 km, in the Lara area, southwest Burgos Province. 67 ichnites of
dinosaurs are documented at Las Sereas 7, identified as theropod and
sauropod trackways occurring in shallow carbonates of lacustrine
environment. Sauropod trackways have intermediate-gauge and low
heteropody, and show different anatomical features to other tracks
found in the ichnological record, especially in the disposition and
orientation of pes digits. They are similar to Polyonyx from the
Middle Jurassic of Portugal. However, since they do not preserve
reliable manus data they are classified as aff. Polyonyx. The three
sauropod trackways are related to the same kind of trackmaker. They
differ from each other only in size, and gregarious behavior has not
been detected. Analysis of these trackways reveals changes in travel
direction even when there are few tracks in each sequence. At the Las
Sereas 7 tracksite, the pace length (PL), width of the angulation
pattern (WAP) and the WAP/PL ratio and depth analysis via
photogrammetry show a direction change in two sauropod trackways. This
tracksite and that at La Pedraja are unique in the
Tithonian-Berriasian interval of the Iberian Peninsula that occur in a
lacustrine environment, and could be indicate of the relationship
between the diversity of Iberian, Tithonian-Berriasian sauropod tracks
and sedimentary environments.


V.F. Santos, P.M. Callapez, D. Castanera, F. Barroso-Barcenilla,
N.P.C. Rodrigues & C.A. Cupeto (2015)
Dinosaur tracks from the Early Cretaceous (Albian) of Parede (Cascais,
Portugal): new contributions for the sauropod palaeobiology of the
Iberian Peninsula.
Journal of Iberian Geology 41(1): 155- 166
doi: http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48662

A recently discovered Early Cretaceous (early late Albian) dinosaur
tracksite at Parede beach (Cascais, Portugal) reveals evidence of
dinoturbation and at least two sauropod trackways. One of these
trackways can be classified as narrow-gauge, which represents unique
evidence in the Albian of the Iberian Peninsula and provides for the
improvement of knowledge of this kind of trackway and its probable
trackmaker, in an age when the sauropod record is scarce. These
dinosaur tracks are preserved on the upper surface of a marly
limestone bed that belongs to the Galé Formation (Água Doce Member,
middle to lower upper Albian). The study of thin-sections of the beds
C22/24 and C26 in the Parede section has revealed a microfacies
composed of foraminifers, radiolarians, ostracods, corals, bivalves,
gastropods, and echinoids in a mainly wackestone texture with
biomicritic matrix. These assemblages match with the lithofacies,
marine molluscs, echinids, and ichnofossils sampled from the section
and indicate a shallow marine, inner shelf palaeoenvironment with a
shallowing-upward trend. The biofacies and the sequence analysis are
compatible with the early late Albian age attributed to the tracksite.
These tracks and the moderate dinoturbation index indicate sauropod
activity in this palaeoenvironment. Titanosaurs can be dismissed as
possible trackmakers on the basis of the narrow-gauge trackway, and
probably by the kidney-shaped manus morphology and the pes-dominated
configuration of the trackway. Narrow-gauge sauropod trackways have
been positively associated with coastal palaeoenvironments, and the
Parede tracksite supports this interpretation. In addition, this
tracksite adds new data about the presence of sauropod pes-dominated
trackways in cohesive substrates. As the Portuguese Cretaceous
sauropod osteological remains are very scarce, the Parede tracksite
yields new and relevant evidence of these dinosaurs. Furthermore, the
Parede tracksite is the youngest evidence of sauropods in the
Portuguese record and some of the rare evidence of sauropods in Europe
during the Albian. This discovery enhances the palaeobiological data
for the Early Cretaceous Sauropoda of the Iberian Peninsula, where the
osteological remains of these dinosaurs are relatively scarce in this
region of southwestern Europe. Therefore, this occurrence is also of
overall interest due to its impact on Cretaceous Sauropoda

I. Díaz-Martínez, E. García-Ortiz & F. Pérez-Lorente (2015)
A new dinosaur tracksite with small footprints in the Urbión Group
(Cameros Basin, Lower Cretaceous, La Rioja, Spain).
Journal of Iberian Geology 41(1): 167-175
doi: http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48661

A new dinosaur tracksite (La Rueda) with ten small tridactyl
footprints (the length ranges between 9 and 15 cm) from the Urbión
Group (Cameros Basin, Lower Cretaceous, La Rioja, Spain) is described.
The footprints are approximately as long as wide and have high
divarication angles between digits II-IV (~80º), some pad impressions
on each digit, claw marks, a medial notch and a circular heel pad
impression. They are here classified as indeterminate ornithopod
footprints and contribute to the increase in the dinosaur
ichnodiversity of the Urbión Group. Small dinosaur footprints are
scarce in the worldwide fossil record. In the Urbión Group, large
dinosaur tracks are much more frequent than small ones. This scarcity
could be explained as ecological biases (dearth of small individuals
in an area). Nevertheless, the number of small footprints in the
Urbión Group is instead the product of by preservation biases (coarse
grain sediments and fluvial erosive bases) and the weathering and
erosion processes (brittle nature of the rock) that affect especially
to small tracks than large ones identified in this Group.