Moussa Masrour, Noura Lkebir & Félix Pérez-Lorente (2017)
Anza palaeoichnological site. Late Cretaceous. Morocco. Part II. Problems of large dinosaur trackways and the first African Macropodosaurus trackway.
Journal of African Earth Sciences(advance online publication)
This area (Anza, Morocco) is an important reference in ichnology because it contains:
The second undebatable African site with pterosaur footprints - described in part I –
The first African site with Macropodosaurus footprints.
The fourth report of this type of footprint in the world.
The first time that the presence of Macropodosaurus has been described in Africa and the second one in which traces attributed to Therizinosaurs in Morocco are mentioned.
The second reported occurrence of pterosaurs tracksites in the Upper Cretaceous of Africa, extending the existence in time of such traces from the Coniacian-Santonian to the Maastrichtian.
The Anza site shows large ichnological surfaces indicating the coexistence in the same area of different vertebrate footprints (dinosaur and pterosaur) and of different types (tridactyl and tetradactyl, semiplantigrade and rounded without digit marks) and the footprint variability of long trackways. This area may become a world reference in ichnology because it contains the second undebatable African site with Cretaceous pterosaur footprints - described in part I - and the first African site with Macropodosaurus footprints. In this work, problems related to long trackways are also analyzed, such as their sinuosity, the order-disorder of the variability (long-short) of the pace length and the difficulty of morphological classification of the theropod footprints due to their morphological variability.