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[dinosaur] Permian–Triassic boundary at Bethulie, Karoo Basin + terminal Permian coproplites from Russian + Permian tetrapod tracks from Tunisia

Ben Creisler

Some recent non-dino papers:

Robert A. Gastaldo, Johann Neveling, John W. Geissman & Jiawen Li (2017)
A multidisciplinary approach to review the vertical and lateral facies relationships of the purported vertebrate-defined terrestrial Permian–Triassic boundary interval at Bethulie, Karoo Basin, South Africa.
Earth-Science Reviews (advance online publication)
doi: https://doi.org/10.1016/j.earscirev.2017.08.002

The stratigraphic section at Bethulie, South Africa, is reported to contain the vertebrate-defined Permian–Triassic boundary succession in the terrestrial realm of the Karoo Basin. The model of vertebrate turnover, from the Daptocephalus to Lystrosaurus Assemblage Zones, tightly constrains the boundary sequence to a short stratigraphic interval where siltstone color begins to change from greenish gray to grayish red, the latter color interpreted to be a consequence of aridification. The biological response to this facies change has been termed “the Great Dying,” and the sedimentary rocks that are preserved are ascribed to a playa lake depositional setting. This drying event is believed to be contemporaneous across the basin, although previous studies have shown that the facies appears at multiple horizons at all purported Permian–Triassic boundary sections in the basin.

Here, we report results of a multidisciplinary effort to characterize the vertebrate assemblage-zone boundary interval exposed at Bethulie using the lithostratigraphic, petrographic, geochemical, and rock magnetic properties of these rocks. In this stratigraphic succession at its “type” location, the 3-m thick assemblage-zone boundary interval is distinguished by thick beds of greenish-gray, greenish-gray mottled to grayish-red, and grayish-red siltstone, all of which change facies characteristics laterally along strike. Specifically, about 220 m to the southeast of the type section, sediments lose all grayish-red coloration, whereas the interval becomes laminated to the northwest. Petrographically, most siltstone is homogenized, with few burrows and small-scale cross-bedded structures with mudchips. There are no gypsiferous or calcareous beds, nor is there evidence of disturbed structures, authigenic breccia, or pseudomorphs associated with dessication. Mean elemental composition of both greenish-gray and grayish-red beds are indistinguishable, geochemically, and both are dominated by illite and chlorite clay species. Mössbauer spectroscopic analyses reveal the presence of a small concentration of fine-grained hematite in the grayish-red siltstone, with its presence mainly found as coatings on clay minerals. Rock magnetic experiments (isothermal remanent magnetization, acquisition and backfield DC demagnetization; magnetic hysteresis; susceptibility vs. temperature) yield data that demonstrate no essential differences between the different colored siltstones. And, both lithologies host mangnetite/maghematite and hematite. Our results do not support the previous interpretation that this inferred Permian–Triassic boundary interval represents the onset of playa lake deposits under conditions of aridification. Rather, our evidence supports the existence of a “wet” landscape at what is considered the Daptocephalus/Lystrosaurus assemblage-zone boundary.


very roughly translating (English version not yet posted)

in Russian:

A. G. Sennikov (2017)
New data on the synecology of the Vyaznikovian terrestrial vertebrate assemblage (terminal Permian, Central Russia).
Paleontological Journal (4): 65-77 

New material is described from the terminal Permian sites of the Vyaznikovian faunal complex from the eastern Vladimir Region, including the tooth of Archosaurus rossicus Tatarinov, 1960, found for the first time in a coprolite. The newly obtained data on tetrapod coprolites allow us to more reliably reconstruct possible food webs and the transformation of the trophic structure of this terrestrial community. The interaction of the higher-order consumers - therocephalians and early archosaurs (proterosuchids) - was not limited to competition for food resources, but was also manifested in the hunting of adults of one taxon on juvenile individuals of another taxon.


Note as well this paper with a free pdf that was posted earlier in preprint on the DML, now in final form:

Piotr Bajdek, Krzysztof Owocki, Andrey G. Sennikov, Valeriy K. Golubev & Grzegorz Niedźwiedzki (2017)
Residues from the Upper Permian carnivore coprolites from Vyazniki in Russia - key questions in reconstruction of feeding habits.
Palaeogeography, Palaeoclimatology, Palaeoecology  482: 70-82
doi: https://doi.org/10.1016/j.palaeo.2017.05.033


Michela Contessi, Sebastian Voigt & Romain Bibonne (2017)
Permian Tetrapod Footprints from Tunisia.
Ichnos (advance online publication)
doi:  http://dx.doi.org/10.1080/10420940.2017.1356724   

Tetrapod footprints have long been known from Permian outcrops of the Cheguimi Sandstone in southern Tunisia, but never described in detail. The only recovered specimen preserves vertebrate tracks of two different biospecies. Ichnomorphotype 1 shows affinity with mid-Permian tetrapod tracks from France (Merifontichnus Gand et al., 2000) and South Africa (unnamed tracks); ichnomorphotype 2 is most similar to Middle to Late Permian tracks of France (Planipes Gand et al., 1995) and Italy (Dicynodontipus Rühle von Lilienstern, 1944), respectively. Both kinds of the Tunisian tetrapod tracks can be referred to non-mammalian therapsid trackmakers. The ichnoassemblage is in agreement with the supposed late Middle to early Late Permian (~Capitanian to Wuchiapingian) age of the Cheguimi Sandstone, although additional finds are necessary to obtain a more precise age control.


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